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CONSOLIDATED STATEMENTS OF CHANGES IN STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Other Comprehensive Loss | Accumulated Deficit
--- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2020 |  | 70,679,190 |  |  |
Beginning balance at Dec. 31, 2020 | $ 296,605 | $ 7 | $ 470,498 | $ (5,010) | $ (168,890)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Net loss | $ (88,641) |  |  |  | (88,641)
Issuance of common stock to directors (in shares) |  | 4,950 |  |  |
Vesting of restricted stock units (in shares) |  | 354,130 |  |  |
Exercise of stock options (in shares) | 423,824 | 423,824 |  |  |
Exercise of stock options | $ 2,786 |  | 2,786 |  |
Stock-based compensation expense | 23,844 |  | 23,844 |  |
Other comprehensive loss | (677) |  |  | (677) |
Ending balance (in shares) at Dec. 31, 2021 |  | 71,462,094 |  |  |
Ending balance at Dec. 31, 2021 | 233,917 | $ 7 | 497,128 | (5,687) | (257,531)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Net loss | $ (150,920) |  |  |  | (150,920)
Issuance of common stock to directors (in shares) |  | 14,928 |  |  |
Vesting of restricted stock units (in shares) |  | 403,648 |  |  |
Exercise of stock options (in shares) | 937,217 | 937,217 |  |  |
Exercise of stock options | $ 25,432 |  | 25,432 |  |
Stock-based compensation expense | 38,830 |  | 38,830 |  |
Other comprehensive loss | (1,559) |  |  | (1,559) |
Ending balance (in shares) at Dec. 31, 2022 |  | 72,817,887 |  |  |
Ending balance at Dec. 31, 2022 | 145,700 | $ 7 | 561,390 | (7,246) | (408,451)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Net loss | (111,441) |  |  |  | (111,441)
Issuance of common stock to directors (in shares) |  | 19,687 |  |  |
Vesting of restricted stock units (in shares) |  | 430,582 |  |  |
Vesting of restricted stock units | $ (9,748) |  | (9,748) |  |
Exercise of stock options (in shares) | 98,610 | 98,610 |  |  |
Exercise of stock options | $ 752 |  | 752 |  |
Stock-based compensation expense | 43,387 |  | 43,387 |  |
Other comprehensive loss | (16,309) |  |  | (16,309) |
Ending balance (in shares) at Dec. 31, 2023 |  | 73,366,766 |  |  |
Ending balance at Dec. 31, 2023 | $ 52,341 | $ 7 | $ 595,781 | $ (23,555) | $ (519,892)

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