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Fair Value Measurements - Schedule of Cash and Available-for-sale Debt Securities' Amortized Cost, Unrealized Gains (Losses) and Fair Value by Significant Investment Category (Details) - USD ($)
Mar. 31, 2020
Dec. 31, 2019
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost $ 109,063,000 $ 138,976,000
Amortized Cost 595,218,000 643,471,000
Unrealized Gain 1,939,000 190,000
Unrealized (Loss) (456,000) (53,000)
Fair Value 596,701,000 643,608,000
Cash    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost 6,398,000 24,631,000
Unrealized Gain 0 0
Unrealized (Loss) 0 0
Fair Value 6,398,000 24,631,000
Money market funds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Unrealized Gain 0 0
Unrealized (Loss) 0 0
Level I | Money market funds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost 62,449,000 32,856,000
Unrealized Gain 0 0
Unrealized (Loss) 0 0
Fair Value 62,449,000 32,856,000
Level II    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost 526,371,000 585,984,000
Unrealized Gain 1,939,000 190,000
Unrealized (Loss) (456,000) (53,000)
Fair Value 527,854,000 586,121,000
Level II | Corporate bonds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost 116,125,000 84,054,000
Unrealized Gain 12,000 22,000
Unrealized (Loss) (456,000) (30,000)
Fair Value 115,681,000 84,046,000
Level II | U.S. treasury securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost 282,330,000 311,083,000
Unrealized Gain 1,831,000 151,000
Unrealized (Loss) 0 (23,000)
Fair Value 284,161,000 311,211,000
Level II | U.S. government agency securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost 55,442,000 95,380,000
Unrealized Gain 96,000 17,000
Unrealized (Loss) 0 0
Fair Value 55,538,000 95,397,000
Level II | Commercial paper    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Amortized Cost 72,474,000 95,467,000
Unrealized Gain 0 0
Unrealized (Loss) 0 0
Fair Value 72,474,000 95,467,000
Cash & Cash Equivalents | Fair Value, Recurring    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 109,063,000 138,976,000
Cash & Cash Equivalents | Fair Value, Recurring | Cash    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 4,211,000 24,631,000
Cash & Cash Equivalents | Fair Value, Recurring | Level I | Money market funds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 55,789,000 26,196,000
Cash & Cash Equivalents | Fair Value, Recurring | Level II    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 49,063,000 88,149,000
Cash & Cash Equivalents | Fair Value, Recurring | Level II | Corporate bonds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Cash & Cash Equivalents | Fair Value, Recurring | Level II | U.S. treasury securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Cash & Cash Equivalents | Fair Value, Recurring | Level II | U.S. government agency securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 10,000,000 22,549,000
Cash & Cash Equivalents | Fair Value, Recurring | Level II | Commercial paper    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 39,063,000 65,600,000
Available-for-sale securities | Fair Value, Recurring    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 478,791,000 497,972,000
Available-for-sale securities | Fair Value, Recurring | Cash    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Available-for-sale securities | Fair Value, Recurring | Level I | Money market funds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Available-for-sale securities | Fair Value, Recurring | Level II    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 478,791,000 497,972,000
Available-for-sale securities | Fair Value, Recurring | Level II | Corporate bonds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 115,681,000 84,046,000
Available-for-sale securities | Fair Value, Recurring | Level II | U.S. treasury securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 284,161,000 311,211,000
Available-for-sale securities | Fair Value, Recurring | Level II | U.S. government agency securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 45,538,000 72,848,000
Available-for-sale securities | Fair Value, Recurring | Level II | Commercial paper    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 33,411,000 29,867,000
Long-term Restricted Cash | Fair Value, Recurring    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 8,847,000 6,660,000
Long-term Restricted Cash | Fair Value, Recurring | Cash    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 2,187,000 0
Long-term Restricted Cash | Fair Value, Recurring | Level I | Money market funds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 6,660,000 6,660,000
Long-term Restricted Cash | Fair Value, Recurring | Level II    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Long-term Restricted Cash | Fair Value, Recurring | Level II | Corporate bonds    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Long-term Restricted Cash | Fair Value, Recurring | Level II | U.S. treasury securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Long-term Restricted Cash | Fair Value, Recurring | Level II | U.S. government agency securities    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value 0 0
Long-term Restricted Cash | Fair Value, Recurring | Level II | Commercial paper    
Fair Value, Balance Sheet Grouping, Financial Statement Captions [Line Items]    
Fair Value $ 0 $ 0