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License agreements - Syros (Details) $ / shares in Units, $ in Thousands, shares in Millions |  |  | 1 Months Ended | 3 Months Ended | 12 Months Ended
--- | --- | --- | --- | --- | ---
Jan. 08, 2018 USD ($) $ / shares | Jan. 01, 2018 USD ($) $ / shares shares | Jan. 31, 2018 USD ($) item $ / shares shares | Mar. 31, 2019 USD ($) | Mar. 31, 2018 USD ($) | Dec. 31, 2018 USD ($)
License agreements |  |  |  |  |  |  |
Long term investment |  |  |  | $ 120,188 |  | $ 99,199 | [1]
Research and development expense |  |  |  | 270,545 | $ 303,103 |  |
Unrealized gain (loss) on long term investments |  |  |  | 20,989 | 22,679 |  |
Syros Pharmaceuticals, Inc. |  |  |  |  |  |  |
License agreements |  |  |  |  |  |  |
Number of program targets | item |  |  | 7 |  |  |  |
Upfront payment under license agreement |  |  | $ 10,000 |  |  |  |
Fair market value of our long term investments |  |  |  | $ 8,600 |  | 5,200 |
Ownership percentage (as a percent) |  |  |  | 3.00% |  |  |
Unrealized gain (loss) on long term investments |  |  |  | $ 3,300 | $ 3,200 |  |
Syros Pharmaceuticals, Inc. | Stock purchase agreement |  |  |  |  |  |  |
License agreements |  |  |  |  |  |  |
Purchase of common stock under Stock Purchase Agreement (in shares ) | shares |  | 0.8 |  |  |  |  |
Purchase price of common stock |  | $ 10,000 |  |  |  |  |
Per share price | $ / shares | $ 9.77 | $ 12.61 |  |  |  |  |
Lock-up period | 12 months |  |  |  |  |  |
Discount for lack of marketability | $ 100 |  |  |  |  |  |
Fair value of shares on the issuance date | $ 7,600 |  |  |  |  |  |
Long term investment |  |  |  |  |  | 7,600 |
Research and development expense |  |  |  |  |  | 2,400 |
Syros Pharmaceuticals, Inc. | Amended stock purchase agreement |  |  |  |  |  |  |
License agreements |  |  |  |  |  |  |
Purchase of common stock under Stock Purchase Agreement (in shares ) | shares |  |  | 0.1 |  |  |  |
Purchase price of common stock |  |  | $ 1,400 |  |  |  |
Per share price | $ / shares |  |  | $ 9.55 |  |  |  |
Long term investment |  |  |  |  |  | $ 1,400 |
Syros Pharmaceuticals, Inc. | Maximum |  |  |  |  |  |  |
License agreements |  |  |  |  |  |  |
Target selection and option exercise fee payment |  |  | $ 54,000 |  |  |  |
Syros Pharmaceuticals, Inc. | Development and Regulatory Milestones | Maximum |  |  |  |  |  |  |
License agreements |  |  |  |  |  |  |
Additional milestone payments under the license agreement |  |  | 50,000 |  |  |  |
Syros Pharmaceuticals, Inc. | Commercialization Milestones | Maximum |  |  |  |  |  |  |
License agreements |  |  |  |  |  |  |
Additional milestone payments under the license agreement |  |  | $ 65,000 |  |  |  |
[1] | The condensed consolidated balance sheet at December 31, 2018 has been derived from the audited financial statements at that date.

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