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License agreements - Syros (Details) $ / shares in Units, $ in Thousands, shares in Millions | 1 Months Ended | 12 Months Ended |  |
--- | --- | --- | --- | ---
Jan. 31, 2018 USD ($) item $ / shares shares | Dec. 31, 2019 USD ($) | Dec. 31, 2018 USD ($) | Dec. 31, 2017 USD ($) | Feb. 28, 2018 USD ($) | Jan. 08, 2018 $ / shares
License agreements |  |  |  |  |  |
Long term investments |  | $ 133,657 | $ 99,199 |  |  |
Research and development expense |  | 1,154,111 | 1,197,957 | $ 1,326,134 |  |
Unrealized gain (loss) on long term investments |  | 34,458 | (44,093) | $ (24,275) |  |
Syros Pharmaceuticals, Inc. |  |  |  |  |  |
License agreements |  |  |  |  |  |
Number of program targets | item | 7 |  |  |  |  |
Upfront payment under license agreement | $ 10,000 |  |  |  |  |
Fair value of shares on the issuance date |  | $ 6,500 | 5,200 |  |  |
Ownership percentage (as a percent) |  | 2.00% |  |  |  |
Unrealized gain (loss) on long term investments |  | $ 1,300 | $ (3,700) |  |  |
Syros Pharmaceuticals, Inc. | Stock purchase agreement |  |  |  |  |  |
License agreements |  |  |  |  |  |
Purchase of common stock under Stock Purchase Agreement (in shares ) | shares | 0.8 |  |  |  |  |
Purchase price of common stock | $ 10,000 |  |  |  |  |
Per share price | $ / shares | $ 12.61 |  |  |  |  | $ 9.77
Lock-up period | 12 months |  |  |  |  |
Discount for lack of marketability | $ 100 |  |  |  |  |
Fair value of shares on the issuance date | 7,600 |  |  |  |  |
Long term investments | 7,600 |  |  |  |  |
Research and development expense | $ 2,400 |  |  |  |  |
Syros Pharmaceuticals, Inc. | Amended stock purchase agreement |  |  |  |  |  |
License agreements |  |  |  |  |  |
Purchase of common stock under Stock Purchase Agreement (in shares ) | shares | 0.1 |  |  |  |  |
Purchase price of common stock | $ 1,400 |  |  |  |  |
Per share price | $ / shares | $ 9.55 |  |  |  |  |
Long term investments |  |  |  |  | $ 1,400 |
Syros Pharmaceuticals, Inc. | Maximum |  |  |  |  |  |
License agreements |  |  |  |  |  |
Target selection and option exercise fee payment | $ 54,000 |  |  |  |  |
Syros Pharmaceuticals, Inc. | Development and Regulatory Milestones | Maximum |  |  |  |  |  |
License agreements |  |  |  |  |  |
Additional milestone payments under the license agreement | 50,000 |  |  |  |  |
Syros Pharmaceuticals, Inc. | Commercialization Milestones | Maximum |  |  |  |  |  |
License agreements |  |  |  |  |  |
Additional milestone payments under the license agreement | $ 65,000 |  |  |  |  |

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