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Property and Equipment, net - Property and equipment, net (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2019	Dec. 31, 2018	Dec. 31, 2017
Property and equipment, net
Property and Equipment, gross	$ 499,437	$ 422,982
Less accumulated depreciation and amortization	(121,870)	(103,231)
Property and Equipment, net	377,567	319,751
Depreciation expense including amortization expense of leasehold improvements	32,100	32,300	$ 24,600
Office equipment
Property and equipment, net
Property and Equipment, gross	15,303	16,955
Laboratory equipment
Property and equipment, net
Property and Equipment, gross	70,510	61,697
Computer equipment
Property and equipment, net
Property and Equipment, gross	59,069	55,436
Land
Property and equipment, net
Property and Equipment, gross	10,203	10,122
Building and leasehold improvements
Property and equipment, net
Property and Equipment, gross	208,293	213,196
Operating lease right-of-use assets
Property and equipment, net
Property and Equipment, gross	19,672
Construction in Progress
Property and equipment, net
Property and Equipment, gross	$ 116,387	$ 65,576

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- Definition

Amount of accumulated depreciation, depletion and amortization for physical assets used in the normal conduct of business to produce goods and services.

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The amount of expense recognized in the current period that reflects the allocation of the cost of tangible assets over the assets' useful lives. Includes production and non-production related depreciation.

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Amount before accumulated depreciation, depletion and amortization of physical assets used in the normal conduct of business and not intended for resale. Examples include, but are not limited to, land, buildings, machinery and equipment, office equipment, and furniture and fixtures.

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