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Property and equipment, net - Property and equipment, net (Details) - USD ($) $ in Thousands | Mar. 31, 2020 | Dec. 31, 2019
--- | --- | ---
Property and equipment, net |  |  |
Property and Equipment, gross | $ 538,366 | $ 499,437 |
Less accumulated depreciation and amortization | (128,332) | (121,870) |
Property and Equipment, net | 410,034 | 377,567 | [1]
Office equipment |  |  |
Property and equipment, net |  |  |
Property and Equipment, gross | 15,285 | 15,303 |
Laboratory equipment |  |  |
Property and equipment, net |  |  |
Property and Equipment, gross | 72,352 | 70,510 |
Computer equipment |  |  |
Property and equipment, net |  |  |
Property and Equipment, gross | 60,077 | 59,069 |
Land |  |  |
Property and equipment, net |  |  |
Property and Equipment, gross | 10,227 | 10,203 |
Building and leasehold improvements |  |  |
Property and equipment, net |  |  |
Property and Equipment, gross | 208,276 | 208,293 |
Operating lease right-of-use assets |  |  |
Property and equipment, net |  |  |
Property and Equipment, gross | 18,641 | 19,672 |
Construction in Progress |  |  |
Property and equipment, net |  |  |
Property and Equipment, gross | $ 153,508 | $ 116,387 |
[1] | The condensed consolidated balance sheet at December 31, 2019 has been derived from the audited consolidated financial statements at that date.

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