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Property and Equipment, net - Property and equipment, net (Details) - USD ($) $ in Thousands	Mar. 31, 2021	Dec. 31, 2020
Property and equipment, net
Property and Equipment, gross	$ 746,075	$ 703,999
Less accumulated depreciation and amortization	(151,405)	(144,374)
Property and Equipment, net	594,670	559,625	[1]
Office equipment
Property and equipment, net
Property and Equipment, gross	18,087	17,880
Laboratory equipment
Property and equipment, net
Property and Equipment, gross	89,714	86,021
Computer equipment
Property and equipment, net
Property and Equipment, gross	67,625	66,640
Land
Property and equipment, net
Property and Equipment, gross	10,328	10,671
Building and leasehold improvements
Property and equipment, net
Property and Equipment, gross	244,872	238,042
Operating lease right-of-use assets
Property and equipment, net
Property and Equipment, gross	25,099	26,816
Construction in Progress
Property and equipment, net
Property and Equipment, gross	$ 290,350	$ 257,929
[1] The condensed consolidated balance sheet at December 31, 2020 has been derived from the audited consolidated financial statements at that date.

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Amount of accumulated depreciation, depletion and amortization for physical assets used in the normal conduct of business to produce goods and services.

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