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License Agreements - Lilly (Details) - USD ($) $ in Thousands	1 Months Ended				3 Months Ended		136 Months Ended
	May 31, 2019	Mar. 31, 2016	Jul. 31, 2010	Dec. 31, 2009	Mar. 31, 2021	Mar. 31, 2020	Mar. 31, 2021
License agreements
Research and Development Expense					$ 306,896	$ 1,085,287
Revenues					604,718	568,507
Product royalty revenues
License agreements
Revenues					$ 99,907	81,780
Eli Lilly
License agreements
Upfront payment received under license agreement				$ 90,000
Eli Lilly | Phase IIB
License agreements
Associated future global development costs from the initiation of a Phase IIb trial, if elected to co-develop (as a percentage)			30.00%		30.00%
Eli Lilly | Pre-specified Events | Maximum
License agreements
Upfront and immediate milestone payment to be received under license agreement				665,000
Eli Lilly | Development Milestones
License agreements
Amount recognized and received for the achievement of a predefined milestone							$ 149,000
Eli Lilly | Development Milestones | Maximum
License agreements
Upfront and immediate milestone payment to be received under license agreement				150,000
Eli Lilly | Regulatory Milestones
License agreements
Amount recognized and received for the achievement of a predefined milestone							$ 265,000
Eli Lilly | Regulatory Milestones | Maximum
License agreements
Upfront and immediate milestone payment to be received under license agreement				365,000
Eli Lilly | Sales and Commercial Milestones
License agreements
Revenues					$ 32,300	25,400
Eli Lilly | Sales and Commercial Milestones | Maximum
License agreements
Upfront and immediate milestone payment to be received under license agreement				$ 150,000
Royalty payments on future global net sales (as a percent)					20.00%
Eli Lilly | Development and Commercialization Milestones
License agreements
Revenues					$ 0	$ 0
Eli Lilly | GVHD
License agreements
Upfront payment under license agreement		$ 35,000
Additional milestone payments under the license agreement		$ 40,000
Eli Lilly | GVHD | Regulatory Milestones
License agreements
Milestone payment made under license agreement	$ 20,000

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