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Collaboration Agreements - Vertex – 2016 Strategic Alliance in Cystic Fibrosis (Details) | 1 Months Ended | 12 Months Ended |
--- | --- | --- | ---
Jul. 31, 2019 USD ($) extension_period | Jul. 31, 2016 USD ($) | Dec. 31, 2020 USD ($) | Dec. 31, 2019 USD ($)
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |
Remaining performance obligations |  |  | $ 87,225,000 |
Increase to the transaction price |  |  | (7,100,000) |
Deferred revenue |  |  | 3,867,193,000 | $ 63,310,000
Additions |  |  | 4,009,960,000 |
Vertex Agreement |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |
Research extension period elected | 6 months |  |  |
Total collaboration revenue | $ 2,000,000.0 |  | 4,500,000 |
Payment received, research extension period | 18 months |  |  |
Number of extension periods | extension_period | 2 |  |  |
Research extension period | 1 year |  |  |
Proceeds from equity investment | $ 20,000,000.0 |  |  |
Termination period | 90 days |  |  |
Termination period upon marketing approval | 180 days |  |  |
Remaining performance obligations |  |  |  | 24,400,000
Increase to the transaction price |  |  | 8,600,000 |
Additions |  | $ 20,000,000.0 |  |
Vertex Agreement | Development Milestones |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |
Estimated arrangement consideration | $ 55,000,000.0 |  |  |
Vertex Agreement | Regulatory Milestones |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |
Estimated arrangement consideration | 220,000,000.0 |  |  |
Vertex Agreement | Regulatory Milestones, Subsequent Products |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |
Estimated arrangement consideration | $ 3,000,000.0 |  |  |
Vertex Agreement | Upfront payment |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |
Remaining performance obligations |  |  | 2,000,000.0 | 20,000,000.0
Vertex Agreement | Research and development |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |
Remaining performance obligations |  |  | $ 2,500,000 | $ 4,400,000

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