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Collaboration Agreements - MERCK Total Transaction Price to the Identified Performance Obligations (Details) - USD ($) $ in Thousands | Dec. 31, 2020 | Dec. 31, 2019 | Jan. 01, 2019
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Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Remaining performance obligations | $ 87,225 |  |
Merck agreement 2015 |  |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Remaining performance obligations | 52,647 | $ 59,735 | $ 65,000
Merck agreement 2015 | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Remaining performance obligations | $ 0 |  |

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- Definition Amount of transaction price allocated to performance obligation that has not been recognized as revenue. + References Reference 1: http://www.xbrl.org/2003/role/disclosureRef -Publisher FASB -Name Accounting Standards Codification -Topic 606 -SubTopic 10 -Section 50 -Paragraph 13 -Subparagraph (a) -URI http://asc.fasb.org/extlink&oid=121604090&loc=SL49130556-203045 + Details
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