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Property and Equipment (Details) - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2020 | Dec. 31, 2019 | Dec. 31, 2018
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | $ 416,550 | $ 296,079 |
Less: Accumulated depreciation | (119,661) | (94,584) |
Property and equipment, net | 296,889 | 201,495 |
Depreciation and amortization | 31,251 | 31,021 | $ 24,862
Laboratory equipment |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | 120,714 | 108,257 |
Leasehold improvements |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | 179,901 | 152,426 |
Furniture, fixtures and other |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | 4,662 | 3,316 |
Computer equipment and software |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | 13,398 | 11,985 |
Internally developed software |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | 7,020 | 7,020 |
Right-of-use asset, financing |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | 56,348 | 9,853 |
Construction in progress |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment, gross | $ 34,507 | $ 3,222 |

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