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Collaboration Agreements - Chiesi Agreement Composition of the Total Transaction Price (Details) - USD ($) $ in Thousands | 1 Months Ended | 12 Months Ended |
--- | --- | --- | ---
Dec. 31, 2020 | Dec. 31, 2020 | Dec. 31, 2019
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Remaining performance obligations | $ 87,225 | $ 87,225 |
Contract with Customer, Liability, Noncurrent | $ 177,419 | 177,419 | $ 138,995
Chiesi agreement |  |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Initial research period | 4 years |  |
Total collaboration revenue |  | $ 200 |
Research extension period elected |  | 1 year |
Research extension period | 4 years |  |
Remaining performance obligations | $ 42,496 | $ 42,496 |
Amounts included in contract liabilities at the beginning of the period |  | 25,000 |
Contract with Customer, Liability, Noncurrent | 25,000 | 25,000 |
Chiesi agreement | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Remaining performance obligations | 17,300 | 17,300 |
Chiesi agreement | Upfront payment |  |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Total collaboration revenue |  | 25,000 |
Remaining performance obligations | 25,000 | $ 25,000 |
Chiesi agreement | Certain Development Regulatory And Commercial Milestone Events |  |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |  |
Amounts included in contract liabilities at the beginning of the period | $ 405,000 |  |

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