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Collaboration Agreements - VEGF Total Transaction Price (Details) - USD ($) $ in Thousands	Dec. 31, 2020	Dec. 31, 2019
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Remaining performance obligations	$ 87,225
VEGF Exercise 2016
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Remaining performance obligations	60,312	$ 57,871
Option exercise fee | VEGF Exercise 2016
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Remaining performance obligations	10,000	10,000
Milestone payment | VEGF Exercise 2016
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Remaining performance obligations	30,000	30,000
Sublicense reimbursement | VEGF Exercise 2016
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Remaining performance obligations	2,250	2,250
Estimated reimbursement for clinical supply | VEGF Exercise 2016
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Remaining performance obligations	$ 18,062	$ 15,621

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Reference 1: http://www.xbrl.org/2003/role/disclosureRef
-Publisher FASB
-Name Accounting Standards Codification
-Topic 606
-SubTopic 10
-Section 50
-Paragraph 13
-Subparagraph (a)
-URI http://asc.fasb.org/extlink&oid=121604090&loc=SL49130556-203045

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