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Collaboration Agreements - VEGF Revenue Recognized (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2020	Dec. 31, 2019	Dec. 31, 2018
VEGF Exercise 2016
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Total collaboration revenue	$ 14,253	$ 583	$ 0

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+ References

No definition available.

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Amount, excluding tax collected from customer, of revenue from satisfaction of performance obligation by transferring promised good or service to customer. Tax collected from customer is tax assessed by governmental authority that is both imposed on and concurrent with specific revenue-producing transaction, including, but not limited to, sales, use, value added and excise.

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