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Collaboration Agreements - VERTEX Composition of the Total Transaction Price (Details)
$ in Thousands
Dec. 31, 2020
USD ($)
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items]  
Remaining performance obligations $ 87,225
Vertex First Extended Research Term  
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items]  
Remaining performance obligations 43,798
Vertex First Extended Research Term | Upfront payment  
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items]  
Remaining performance obligations 4,000
Vertex First Extended Research Term | Research and development  
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items]  
Remaining performance obligations $ 39,798