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CONSOLIDATED STATEMENTS OF INCOME (UNAUDITED) - USD ($) shares in Thousands, $ in Thousands | 3 Months Ended
--- | ---
Mar. 31, 2022 | Mar. 31, 2021
Interest income |  |
Interest and fees on loans | $ 285,698 | $ 274,100
Mortgage loans held-for-sale | 6,087 | 9,036
Interest-bearing deposits with banks | 1,687 | 1,199
Federal funds sold and securities purchased under resale agreements | 431 | 0
Investment securities | 32,398 | 19,264
Trading account securities | 5 | 2
Federal Home Loan Bank and Federal Reserve Bank stock | 1,772 | 1,745
Brokerage customer receivables | 174 | 123
Total interest income | 328,252 | 305,469
Interest expense |  |
Interest on deposits | 14,854 | 27,944
Interest on Federal Home Loan Bank advances | 4,816 | 4,840
Interest on other borrowings | 2,239 | 2,609
Interest on subordinated notes | 5,482 | 5,477
Interest on junior subordinated debentures | 1,567 | 2,704
Total interest expense | 28,958 | 43,574
Net interest income | 299,294 | 261,895
Provision for credit losses | 4,106 | (45,347)
Net interest income after provision for credit losses | 295,188 | 307,242
Non-interest income |  |
Revenue from contracts with customers | 54,552 | 47,768
Mortgage banking | 77,231 | 113,494
(Losses) gains on investment securities, net | (2,782) | 1,154
Fees from covered call options | 3,742 | 0
Trading gains, net | 3,889 | 419
Operating lease income, net | 15,475 | 14,440
Other | 18,558 | 15,654
Total non-interest income | 162,790 | 186,506
Non-interest expense |  |
Salaries and employee benefits | 172,355 | 180,809
Software and equipment | 22,810 | 20,912
Operating lease equipment depreciation | 9,708 | 10,771
Occupancy, net | 17,824 | 19,996
Data processing | 7,505 | 6,048
Advertising and marketing | 11,924 | 8,546
Professional fees | 8,401 | 7,587
Amortization of other acquisition-related intangible assets | 1,609 | 2,007
FDIC insurance | 7,729 | 6,558
Other real estate owned expense, net | (1,032) | (251)
Other | 25,465 | 23,906
Total non-interest expense | 284,298 | 286,889
Income before taxes | 173,680 | 206,859
Income tax expense | 46,289 | 53,711
Net income | 127,391 | 153,148
Preferred stock dividends | 6,991 | 6,991
Net income applicable to common shares | $ 120,400 | $ 146,157
Net income per common share-Basic (usd per share) | $ 2.11 | $ 2.57
Net income per common share-Diluted (usd per share) | 2.07 | 2.54
Cash dividends declared per common share (usd per share) | $ 0.34 | $ 0.31
Weighted average common shares outstanding (in shares) | 57,196 | 56,904
Dilutive potential common shares (in shares) | 862 | 681
Average common shares and dilutive common shares (in shares) | 58,058 | 57,585
Wealth management |  |
Non-interest income |  |
Revenue from contracts with customers | $ 31,394 | $ 29,309
Service charges on deposit accounts |  |
Non-interest income |  |
Revenue from contracts with customers | $ 15,283 | $ 12,036

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