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CONSOLIDATED STATEMENTS OF CONDITION - USD ($) $ in Thousands | Mar. 31, 2022 | Dec. 31, 2021 | Mar. 31, 2021
--- | --- | --- | ---
Assets |  |  |
Cash and due from banks | $ 462,516 | $ 411,150 | $ 426,325
Federal funds sold and securities purchased under resale agreements | 700,056 | 700,055 | 52
Interest-bearing deposits with banks | 4,013,597 | 5,372,603 | 3,348,794
Available-for-sale securities, at fair value | 2,998,898 | 2,327,793 | 2,430,749
Held-to-maturity securities, at amortized cost, net of allowance for credit losses of $159, $78 and $99 at March 31, 2022, December 31, 2021 and March 31, 2021, respectively ($3.1 billion, $2.9 billion and $2.1 billion fair value at March 31, 2022, December 31, 2021 and March 31, 2021, respectively) | 3,435,729 | 2,942,285 | 2,166,419
Trading account securities | 852 | 1,061 | 951
Equity securities with readily determinable fair value | 92,689 | 90,511 | 90,338
Federal Home Loan Bank and Federal Reserve Bank stock | 136,163 | 135,378 | 135,881
Brokerage customer receivables | 22,888 | 26,068 | 19,056
Mortgage loans held-for-sale, at fair value | 606,545 | 817,912 | 1,260,193
Loans, net of unearned income | 35,280,547 | 34,789,104 | 33,171,233
Allowance for loan losses | (301,168) | (299,653) | (321,209)
Premises, software and equipment, net | 761,213 | 766,405 | 760,522
Lease investments, net | 240,656 | 242,082 | 238,984
Accrued interest receivable and other assets | 1,066,750 | 1,084,115 | 1,230,362
Goodwill | 655,402 | 655,149 | 646,017
Other acquisition-related intangible assets | 26,699 | 28,307 | 34,035
Total assets | 50,250,661 | 50,142,143 | 45,682,202
Deposits: |  |  |
Non-interest-bearing | 13,748,918 | 14,179,980 | 12,297,337
Interest-bearing | 28,470,404 | 27,915,605 | 25,575,315
Total deposits | 42,219,322 | 42,095,585 | 37,872,652
Federal Home Loan Bank advances | 1,241,071 | 1,241,071 | 1,228,436
Other borrowings | 482,516 | 494,136 | 516,877
Subordinated notes | 437,033 | 436,938 | 436,595
Junior Subordinated Debentures | 253,566 | 253,566 | 253,566
Trade date securities payable | 437 | 0 | 995
Accrued interest payable and other liabilities | 1,124,460 | 1,122,159 | 1,120,570
Total liabilities | 45,758,405 | 45,643,455 | 41,429,691
Shareholders’ Equity: |  |  |
Common stock, no par value; $1.00 stated value; 100,000,000 shares authorized at March 31, 2022, December 31, 2021 and March 31, 2021; 59,090,903 shares issued at March 31, 2022, 58,891,780 shares issued at December 31, 2021 and 58,726,900 shares issued at March 31, 2021 | 59,091 | 58,892 | 58,727
Surplus | 1,698,093 | 1,685,572 | 1,663,008
Treasury stock, at cost, 1,837,689 shares at March 31, 2022, 1,837,689 shares at December 31, 2021, and 1,703,627 shares at March 31, 2021 | (109,903) | (109,903) | (100,363)
Retained earnings | 2,548,474 | 2,447,535 | 2,208,535
Accumulated other comprehensive (loss) income | (115,999) | 4,092 | 10,104
Total shareholders’ equity | 4,492,256 | 4,498,688 | 4,252,511
Total liabilities and shareholders’ equity | 50,250,661 | 50,142,143 | 45,682,202
Loans |  |  |
Assets |  |  |
Loans, net of unearned income | 35,280,547 | 34,789,104 | 33,171,233
Allowance for loan losses | (250,539) | (247,835) | (277,709)
Net loans | 35,030,008 | 34,541,269 | 32,893,524
Series D - $25 liquidation value; 5,000,000 shares issued and outstanding at March 31, 2022, December 31, 2021 and March 31, 2021 |  |  |
Shareholders’ Equity: |  |  |
Preferred stock, no par value; 20,000,000 shares authorized: | 125,000 | 125,000 | 125,000
Series E - $25,000 liquidation value; 11,500 shares issued and outstanding at March 31, 2022, December 31, 2021 and March 31, 2021 |  |  |
Shareholders’ Equity: |  |  |
Preferred stock, no par value; 20,000,000 shares authorized: | $ 287,500 | $ 287,500 | $ 287,500

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