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Income Taxes - Schedule of Significant Components of Net Deferred Tax Assets (Liabilities) (Details) - USD ($) $ in Thousands | Sep. 30, 2021 | Dec. 31, 2020
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Deferred Tax Assets |  |
Allowance for credit losses, net of recapture | $ 28,791 | $ 34,971
Benefit plan accruals | 15,579 | 20,076
Net operating loss carryforwards | 14,823 | 18,982
Deferred gain on securities | 1,334 | 2,102
Acquired loans | 8,949 | 11,989
Operating lease liabilities | 22,959 | 24,245
Tax credit investments and other partnerships | 494 | 1,054
Other, net | 942 | 488
Total deferred tax assets | 93,871 | 113,907
Deferred Tax Liabilities |  |
Purchase accounting | (18,524) | (18,232)
Loan servicing rights | (7,179) | (6,582)
Premises and equipment | (14,550) | (14,008)
Prepaid expenses | (953) | (955)
Operating lease right-of-use assets | (21,106) | (21,569)
Unrealized gains on available-for-sale investment securities | (14,590) | (40,756)
Unrealized gains on hedges | (628) | (1,080)
Other, net | (1,561) | (1,555)
Total deferred tax liabilities | (79,091) | (104,737)
Net deferred tax assets | $ 14,780 | $ 9,170

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