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Income Taxes - Deferred Tax Assets and Liabilities (Details) - USD ($) $ in Thousands | Dec. 31, 2021 | Dec. 31, 2020
--- | --- | ---
Deferred income tax assets |  |
Allowance for loan losses | $ 112,180 | $ 165,691
Lease liability | 102,412 | 98,340
Employee benefits and deferred compensation | 35,782 | 27,917
Net operating loss carryforwards | 23,897 | 29,684
Net unrealized losses on investment securities available for sale and cash flow hedges | 19,261 | 0
Deferred revenue | 18,487 | 24,751
Tax credit carryforwards | 15,870 | 8,605
Fair value of investment securities and loans | 6,439 | 10,093
Non-performing loan interest | 2,621 | 12,472
Other | 7,100 | 9,819
Deferred Tax Asset, Before Valuation Allowance | 344,049 | 387,372
Less valuation allowance | (19,003) | (19,191)
Total deferred tax assets | 325,046 | 368,181
Deferred tax liabilities |  |
Right-of-use asset | (98,773) | (98,681)
Excess tax over financial statement depreciation | (27,355) | (40,452)
Purchase accounting intangibles | (14,820) | (14,458)
Other properties held for sale | (5,345) | (3,025)
Prepaid expense | (4,631) | (5,955)
Net unrealized gains on investment securities available for sale and cash flow hedges | 0 | (64,344)
Other | (5,071) | (10,418)
Total gross deferred tax liabilities | (155,995) | (237,333)
Net deferred tax assets | $ 169,051 | $ 130,848

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