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Selected quarterly financial data (Details) - USD ($) $ / shares in Units, $ in Thousands | 3 Months Ended | 12 Months Ended
--- | --- | ---
Dec. 31, 2015 | Sep. 30, 2015 | Jun. 30, 2015 | Mar. 31, 2015 | Dec. 31, 2014 | Sep. 30, 2014 | Jun. 30, 2014 | Mar. 31, 2014 | Dec. 31, 2015 | Dec. 31, 2014 | Dec. 31, 2013
Selected quarterly financial data (Unaudited) |  |  |  |  |  |  |  |  |  |  |
Net product revenue | $ 12,694 | $ 9,772 | $ 6,161 | $ 5,069 | $ 636 | $ 81 |  |  | $ 33,696 | $ 717 |
Collaboration and grant revenue | 40 | 4 | 613 | 2,413 | 12,022 | 1,613 | $ 1,677 | $ 9,217 | 3,070 | 24,528 | $ 34,696
Operating expenses | 60,935 | 52,008 | 45,400 | 45,553 | 44,888 | 29,295 | 27,046 | 23,429 | 203,896 | 124,658 | 80,094
Loss from operations | (48,201) | (42,232) | (38,626) | (38,071) | (32,230) | (27,601) | (25,369) | (14,212) | (167,130) | (99,413) | (45,398)
Net loss | $ (50,948) | $ (43,223) | $ (38,361) | $ (37,915) | $ (27,269) | $ (27,282) | $ (25,104) | $ (14,098) | (170,447) | (93,753) | (51,574)
Deemed dividend |  |  |  |  |  |  |  |  |  |  | 18,249
Gain on exchange of convertible preferred stock in connection with recapitalization |  |  |  |  |  |  |  |  |  |  | 3,391
Net loss attributable to common stockholders |  |  |  |  |  |  |  |  | $ (170,447) | $ (93,753) | $ (66,432)
Net loss per share-basic and diluted (in dollars per share) | $ (1.50) | $ (1.27) | $ (1.14) | $ (1.15) | $ (0.84) | $ (0.93) | $ (0.86) | $ (0.58) | $ (5.07) | $ (2.97) | $ (5.18)

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