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v3.3.1.900

Collaborations and grants (Details) $ in Millions |  | 1 Months Ended | 12 Months Ended
--- | --- | --- | ---
Jan. 02, 2011 item | Nov. 30, 2014 USD ($) | Jan. 31, 2014 USD ($) | Nov. 30, 2011 USD ($) item | Dec. 31, 2015 USD ($) | Dec. 31, 2014 USD ($) | Dec. 31, 2013 USD ($)
Minimum |  |  |  |  |  |  |
Grant revenue |  |  |  |  |  |  |
Term of grant program |  |  |  |  | 2 years |  |
Maximum |  |  |  |  |  |  |
Grant revenue |  |  |  |  |  |  |
Term of grant program |  |  |  |  | 5 years |  |
Collaboration agreement | Development event milestones |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Collaboration revenue recognized |  | $ 10.0 | $ 7.5 |  |  |  |
Collaboration agreement | Roche and SMA Foundation [Member] |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Number of compounds in preclinical development | item |  |  |  | 3 |  |  |
Deferred revenue |  |  |  | $ 30.0 |  |  |
Number of significant deliverables | item |  |  |  | 2 |  |  |
Research period for applying technology |  |  |  | 2 years |  |  |
Collaboration revenue recognized |  |  |  |  | $ 0.6 | $ 20.2 | $ 26.6
Collaboration agreement | Roche and SMA Foundation [Member] | Development event milestones | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | 135.0 |  |
Collaboration agreement | Roche and SMA Foundation [Member] | Sales milestones | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | 325.0 |  |
Early stage collaboration and discovery agreements |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Deferred revenue |  |  |  |  | 0.0 | 1.8 |
Number of significant deliverables | item | 2 |  |  |  |  |  |
Collaboration revenue recognized |  |  |  |  | $ 1.8 | $ 1.3 |
Early stage collaboration and discovery agreements | Minimum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Research period for applying technology |  |  |  |  | 3 years |  |
Early stage collaboration and discovery agreements | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Research period for applying technology |  |  |  |  | 4 years |  |
Early stage collaboration and discovery agreements | Development event milestones |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | $ 143.0 |  |
Early stage collaboration and discovery agreements | Sales milestones | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | $ 252.0 |  |

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