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Collaborations and grants (Details) |  | 1 Months Ended | 12 Months Ended
--- | --- | --- | ---
Jan. 02, 2011 item | Nov. 30, 2014 USD ($) | Jan. 31, 2014 USD ($) | Nov. 30, 2011 USD ($) item | Dec. 31, 2016 USD ($) | Dec. 31, 2015 USD ($) | Dec. 31, 2014 USD ($)
Minimum |  |  |  |  |  |  |
Grant revenue |  |  |  |  |  |  |
Term of grant program |  |  |  |  | 2 years |  |
Maximum |  |  |  |  |  |  |
Grant revenue |  |  |  |  |  |  |
Term of grant program |  |  |  |  | 5 years |  |
Collaboration agreement | Development event milestones |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Collaboration revenue recognized |  | $ 10,000,000 | $ 7,500,000 |  |  |  |
Collaboration agreement | Roche and SMA Foundation [Member] |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Number of compounds in preclinical development | item |  |  |  | 3 |  |  |
Deferred revenue |  |  |  | $ 30,000,000 |  |  |
Number of significant deliverables | item |  |  |  | 2 |  |  |
Research period for applying technology |  |  |  | 2 years |  |  |
Collaboration revenue recognized |  |  |  |  | $ 400,000 | $ 600,000 | $ 20,200,000
Collaboration agreement | Roche and SMA Foundation [Member] | Development event milestones | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | 135,000,000 |  |
Collaboration agreement | Roche and SMA Foundation [Member] | Sales milestones | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | 325,000,000 |  |
Early stage collaboration and discovery agreements |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Deferred revenue |  |  |  |  | $ 0 | 0 | 1,800,000
Number of significant deliverables | item | 2 |  |  |  |  |  |
Collaboration revenue recognized |  |  |  |  |  | $ 1,800,000 | $ 1,300,000
Early stage collaboration and discovery agreements | Minimum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Research period for applying technology |  |  |  |  | 3 years |  |
Early stage collaboration and discovery agreements | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Research period for applying technology |  |  |  |  | 4 years |  |
Early stage collaboration and discovery agreements | Development event milestones |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | $ 143,000,000 |  |
Early stage collaboration and discovery agreements | Sales milestones | Maximum |  |  |  |  |  |  |
Collaborations and grants |  |  |  |  |  |  |
Additional consideration receivable upon achievement of specified events |  |  |  |  | $ 252,000,000 |  |

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- Definition Revenue from multiple-deliverable arrangements that include licensing fees and services revenue. Licensing revenue is consideration received from another party for the right to use, but not own, certain of the entity's intangible assets. Licensing arrangements include, but are not limited to, rights to use a patent, copyright, technology, manufacturing process, software or trademark. Licensing fees are generally, but not always, fixed as to amount and not dependent upon the revenue generated by the licensing party. An entity may receive licensing fees for licenses that also generate royalty payments to the entity. Services revenue may be derived by providing other, nonspecified, services during the reporting period. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 225 -SubTopic 10 -Section S99 -Paragraph 2 -Subparagraph (SX 210.5-03.1(d),(e)) -URI http://asc.fasb.org/extlink&oid=63488584&loc=d3e20235-122688 + Details
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