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Selected quarterly financial data (Details) - USD ($) $ / shares in Units, $ in Thousands | 3 Months Ended | 12 Months Ended
--- | --- | ---
Dec. 31, 2019 | Sep. 30, 2019 | Jun. 30, 2019 | Mar. 31, 2019 | Dec. 31, 2018 | Sep. 30, 2018 | Jun. 30, 2018 | Mar. 31, 2018 | Dec. 31, 2019 | Dec. 31, 2018 | Dec. 31, 2017
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |  |  | $ 306,980 | $ 264,734 | $ 194,392
Operating expenses | $ 169,243 | $ 131,891 | $ 124,280 | $ 122,723 | $ 131,476 | $ 101,821 | $ 74,317 | $ 72,805 | 548,138 | 380,419 | 258,684
Loss from operations | (72,784) | (60,475) | (38,758) | (69,140) | (45,138) | (48,230) | (5,574) | (16,743) | (241,158) | (115,685) | (64,292)
Net loss attributable to common stockholders | $ (77,677) | $ (59,997) | $ (41,789) | $ (72,113) | $ (48,330) | $ (50,969) | $ (9,520) | $ (19,263) | $ (251,576) | $ (128,081) | $ (79,000)
Basic and diluted net loss per common share (in dollars per share) | $ (1.37) | $ (1.06) | $ (0.75) | $ (1.29) | $ 0.96 | $ (1.06) | $ (0.21) | $ (0.46) | $ (4.27) | $ (2.75) | $ (2.02)
Net product revenue |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Revenue | $ 81,407 | $ 71,369 | $ 85,476 | $ 53,054 | $ 85,833 | $ 53,021 | $ 68,170 | $ 55,981 | $ 291,306 | $ 263,005 | $ 174,066
Collaboration and grant revenue |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Revenue | $ 15,052 | $ 47 | $ 46 | $ 529 | $ 505 | $ 570 | $ 573 | $ 81 | $ 15,674 | $ 1,729 | $ 20,326

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