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Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Common stock | Additional paid-in capital Cumulative Effect Adjustment | Additional paid-in capital | Accumulated other comprehensive income (loss) | Accumulated deficit Cumulative Effect Adjustment | Accumulated deficit | Cumulative Effect Adjustment | Total
--- | --- | --- | --- | --- | --- | --- | --- | ---
Balance (in shares) at Dec. 31, 2019 | 61,935,870 |  |  |  |  |  |  |
Balance at the beginning of the period at Dec. 31, 2019 | $ 62 |  | $ 1,795,351 | $ (10,584) |  | $ (1,190,499) |  | $ 594,330
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Issuance of common stock related to equity offerings (in shares) | 368,514 |  |  |  |  |  |  |
Issuance of common stock related to equity offering |  |  | 18,950 |  |  |  |  | 18,950
Issuance of common stock related to acquisition (in shares) | 845,364 |  |  |  |  |  |  |
Issuance of common stock related to acquisition | $ 1 |  | 42,868 |  |  |  |  | 42,869
Issuance of common stock related to rights exchange (in shares) | 2,821,176 |  |  |  |  |  |  |
Issuance of common stock related to rights exchange | $ 3 |  | 150,525 |  |  |  |  | 150,528
Exercise of options (in shares) | 1,034,288 |  |  |  |  |  |  |
Exercise of options | $ 1 |  | 25,044 |  |  |  |  | 25,045
Restricted stock vesting and issuance (in shares) | 176,776 |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan (in shares) | 58,691 |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan |  |  | 2,406 |  |  |  |  | 2,406
Share-based compensation expense |  |  | 32,130 |  |  |  |  | 32,130
Other |  |  |  |  |  | (218) |  | (218)
Net loss |  |  |  |  |  | (294,114) |  | (294,114)
Comprehensive income (loss) |  |  |  | 568 |  |  |  | 568
Balance (in shares) at Jun. 30, 2020 | 67,240,679 |  |  |  |  |  |  |
Balance at the end of the period at Jun. 30, 2020 | $ 67 |  | 2,067,274 | (10,016) |  | (1,484,831) |  | 572,494
Balance (in shares) at Mar. 31, 2020 | 62,758,520 |  |  |  |  |  |  |
Balance at the beginning of the period at Mar. 31, 2020 | $ 62 |  | 1,834,061 | (1,985) |  | (1,303,404) |  | 528,734
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Issuance of common stock related to equity offerings (in shares) | 106,309 |  |  |  |  |  |  |
Issuance of common stock related to equity offering |  |  | 5,447 |  |  |  |  | 5,447
Issuance of common stock related to acquisition (in shares) | 845,364 |  |  |  |  |  |  |
Issuance of common stock related to acquisition | $ 1 |  | 42,868 |  |  |  |  | 42,869
Issuance of common stock related to rights exchange (in shares) | 2,821,176 |  |  |  |  |  |  |
Issuance of common stock related to rights exchange | $ 3 |  | 150,525 |  |  |  |  | 150,528
Exercise of options (in shares) | 654,604 |  |  |  |  |  |  |
Exercise of options | $ 1 |  | 15,057 |  |  |  |  | 15,058
Restricted stock vesting and issuance (in shares) | (3,985) |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan (in shares) | 58,691 |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan |  |  | 2,406 |  |  |  |  | 2,406
Share-based compensation expense |  |  | 16,910 |  |  |  |  | 16,910
Net loss |  |  |  |  |  | (181,427) |  | (181,427)
Comprehensive income (loss) |  |  |  | (8,031) |  |  |  | (8,031)
Balance (in shares) at Jun. 30, 2020 | 67,240,679 |  |  |  |  |  |  |
Balance at the end of the period at Jun. 30, 2020 | $ 67 |  | 2,067,274 | (10,016) |  | (1,484,831) |  | 572,494
Balance (in shares) at Dec. 31, 2020 | 69,718,096 |  |  |  |  |  |  |
Balance at the beginning of the period at Dec. 31, 2020 | $ 70 | $ (175,236) | 2,171,746 | (60,957) | $ 54,796 | (1,628,877) | $ (120,440) | 481,982
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Exercise of options (in shares) | 475,942 |  |  |  |  |  |  |
Exercise of options |  |  | 13,144 |  |  |  |  | 13,144
Restricted stock vesting and issuance (in shares) | 292,171 |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan (in shares) | 73,121 |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan |  |  | 2,627 |  |  |  |  | 2,627
Share-based compensation expense |  |  | 51,406 |  |  |  |  | 51,406
Net loss |  |  |  |  |  | (247,012) |  | (247,012)
Comprehensive income (loss) |  |  |  | 14,870 |  |  |  | 14,870
Balance (in shares) at Jun. 30, 2021 | 70,559,330 |  |  |  |  |  |  |
Balance at the end of the period at Jun. 30, 2021 | $ 70 |  | 2,063,687 | (46,087) |  | (1,821,093) |  | 196,577
Balance (in shares) at Mar. 31, 2021 | 70,405,905 |  |  |  |  |  |  |
Balance at the beginning of the period at Mar. 31, 2021 | $ 70 |  | 2,033,972 | (38,743) |  | (1,702,723) |  | 292,576
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Exercise of options (in shares) | 60,159 |  |  |  |  |  |  |
Exercise of options |  |  | 1,389 |  |  |  |  | 1,389
Restricted stock vesting and issuance (in shares) | 20,145 |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan (in shares) | 73,121 |  |  |  |  |  |  |
Issuance of common stock in connection with an employee stock purchase plan |  |  | 2,627 |  |  |  |  | 2,627
Share-based compensation expense |  |  | 25,699 |  |  |  |  | 25,699
Net loss |  |  |  |  |  | (118,370) |  | (118,370)
Comprehensive income (loss) |  |  |  | (7,344) |  |  |  | (7,344)
Balance (in shares) at Jun. 30, 2021 | 70,559,330 |  |  |  |  |  |  |
Balance at the end of the period at Jun. 30, 2021 | $ 70 |  | $ 2,063,687 | $ (46,087) |  | $ (1,821,093) |  | $ 196,577

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