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Revenue recognition - Collaboration and Royalty Revenue (Details) - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2022	Mar. 31, 2021	Nov. 30, 2011
Collaboration and royalty revenue
Remaining potential milestones that can be achieved	$ 300,000
Revenue	148,735	$ 117,942
Collaboration revenue
Collaboration and royalty revenue
Revenue	7	20,007
Royalty revenue
Collaboration and royalty revenue
Revenue	18,896	6,655
SMA License Agreement | Collaboration revenue
Collaboration and royalty revenue
Revenue	0	20,000
SMA License Agreement | Royalty revenue
Collaboration and royalty revenue
Revenue	18,900	6,700
SMA License Agreement | Roche And Sma Foundation | Collaboration revenue
Collaboration and royalty revenue
Revenue	0	20,000
Research And Development Event Milestones | SMA License Agreement
Collaboration and royalty revenue
Milestone payments received		20,000
Sales Milestones | SMA License Agreement
Collaboration and royalty revenue
Remaining potential milestones that can be achieved	$ 300,000
Milestone payments received		$ 20,000
Maximum | Research And Development Event Milestones | SMA License Agreement
Collaboration and royalty revenue
Remaining potential milestones that can be achieved			$ 135,000
Maximum | Sales Milestones | SMA License Agreement
Collaboration and royalty revenue
Remaining potential milestones that can be achieved			$ 325,000

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