Balance at

Beginning of

End of

Nine Months Ended September 30, 2021

Period

Additions

Deductions

Period

Contract assets:

Receivable from collaboration partner - related party

$

2,426

$

5,732

$

(5,520)

$

2,638

Contract liabilities:

Deferred revenue - related party

$

14,477

$

16,715

$

(28,951)

$

2,241

Payable to collaboration partner - related party

$

2,732

$

9,785

$

(11,396)

$

1,121

Balance at

Beginning of

End of

Nine Months Ended September 30, 2020

Period

Additions

Deductions

Period

Contract assets:

Receivable from collaboration partner - related party

$

5,955

$

5,284

$

(8,450)

$

2,789

Contract asset - related party

$

800

$

342

$

(1,142)

$

—

Contract liabilities:

Deferred revenue - related party

$

41,530

$

3,500

$

(24,153)

$

20,877

Payable to collaboration partner - related party

$

1,262

$

2,114

$

(1,299)

$

2,077