Balance at

Beginning of

End of

Three Months Ended March 31, 2022

Period

Additions

Deductions

Period

Contract assets:

Receivable from collaboration partner - related party

$

1,566

$

25,150

$

(1,566)

$

25,150

Contract liabilities:

Deferred revenue - related party

$

1,601

$

25,658

$

(26,491)

$

768

Payable to collaboration partner - related party

$

899

$

330

$

(849)

$

380

Balance at

Beginning of

End of

Three Months Ended March 31, 2021

Period

Additions

Deductions

Period

Contract assets:

Receivable from collaboration partner - related party

$

2,426

$

1,570

$

—

$

3,996

Contract liabilities:

Deferred revenue - related party

$

14,477

$

1,017

$

(9,726)

$

5,768

Payable to collaboration partner - related party

$

2,732

$

4,091

$

—

$

6,823