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Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-in Capital | Accumulated Deficit | Accumulated Other Comprehensive Income (Loss)
--- | --- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2014 | $ 309,489 | $ 10 | $ 807,631 | $ (498,143) | $ (9)
Beginning balance (in shares) at Dec. 31, 2014 |  | 100,047,331 |  |  |
Issuance of common stock from exercise of stock options | 12,991 |  | 12,991 |  |
Issuance of common stock from exercise of stock options (in shares) |  | 1,822,578 |  |  |
Issuance of common stock pursuant to employee stock purchase plan | 1,556 |  | 1,556 |  |
Issuance of common stock pursuant to employee stock purchase plan (in shares) |  | 68,793 |  |  |
Income tax benefit from exercise of stock options | 247 |  | 247 |  |
Deferred offering costs | (292) |  | (292) |  |
Net loss | (164,443) |  |  | (164,443) |
Stock-based compensation | 40,194 |  | 40,194 |  |
Other comprehensive income (loss) | 20 |  |  |  | 20
Ending balance at Dec. 31, 2015 | 199,762 | $ 10 | 862,327 | (662,586) | 11
Ending balance (in shares) at Dec. 31, 2015 |  | 101,938,702 |  |  |
Issuance of common stock in public offering, net of issuance costs | 497,765 | $ 2 | 497,763 |  |
Issuance of common stock in public offering, net of issuance costs (in shares) |  | 17,314,523 |  |  |
Issuance of common stock from exercise of stock options | 18,000 |  | 18,000 |  |
Issuance of common stock from exercise of stock options (in shares) |  | 1,977,661 |  |  |
Issuance of common stock pursuant to employee stock purchase plan | 3,131 |  | 3,131 |  |
Issuance of common stock pursuant to employee stock purchase plan (in shares) |  | 136,283 |  |  |
Income tax benefit from exercise of stock options | 596 |  | 596 |  |
Proceeds from settlement agreement | 14,320 |  | 14,320 |  |
Net loss | (271,393) |  |  | (271,393) |
Stock-based compensation | 56,135 |  | 56,135 |  |
Other comprehensive income (loss) | 95 |  |  |  | 95
Ending balance at Dec. 31, 2016 | 518,411 | $ 12 | 1,452,272 | (933,979) | 106
Ending balance (in shares) at Dec. 31, 2016 |  | 121,367,169 |  |  |
Issuance of common stock from exercise of stock options | $ 26,665 |  | 26,665 |  |
Issuance of common stock from exercise of stock options (in shares) | 1,442,411 | 1,442,411 |  |  |
Issuance of common stock pursuant to employee stock purchase plan | $ 4,522 |  | 4,522 |  |
Issuance of common stock pursuant to employee stock purchase plan (in shares) |  | 192,402 |  |  |
Issuance of common stock from exercise of warrants on a net issuance basis (in shares) | 1,408,570 | 1,408,570 |  |  |
Net loss | $ (289,403) |  |  | (289,403) |
Cumulative effect adjustment from adoption of ASU 2016-09 | (289) |  |  | (289) |
Stock-based compensation | 75,884 |  | 75,884 |  |
Other comprehensive income (loss) | (505) |  |  |  | (505)
Ending balance at Dec. 31, 2017 | $ 335,285 | $ 12 | $ 1,559,343 | $ (1,223,671) | $ (399)
Ending balance (in shares) at Dec. 31, 2017 |  | 124,410,552 |  |  |

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