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Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-in Capital | Accumulated Deficit | Accumulated Other Comprehensive Income (Loss)
--- | --- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2016 | $ 518,411 | $ 12 | $ 1,452,272 | $ (933,979) | $ 106
Beginning balance (in shares) at Dec. 31, 2016 |  | 121,367,169 |  |  |
Issuance of common stock from exercise of stock options | 26,665 |  | 26,665 |  |
Issuance of common stock from exercise of stock options (in shares) |  | 1,442,411 |  |  |
Issuance of common stock pursuant to employee stock purchase plan | 4,522 |  | 4,522 |  |
Issuance of common stock pursuant to employee stock purchase plan (in shares) |  | 192,402 |  |  |
Issuance of common stock from exercise of warrants on a net issuance basis (in shares) |  | 1,408,570 |  |  |
Net loss | (289,403) |  |  | (289,403) |
Cumulative effect adjustment from adoption of ASU 2016-09 | (289) |  |  | (289) |
Stock-based compensation | 75,884 |  | 75,884 |  |
Other comprehensive income (loss) | (505) |  |  |  | (505)
Ending balance at Dec. 31, 2017 | 335,285 | $ 12 | 1,559,343 | (1,223,671) | (399)
Beginning balance (in shares) at Dec. 31, 2017 |  | 124,410,552 |  |  |
Issuance of common stock in public offering, net of issuance costs | 298,537 | $ 2 | 298,535 |  |
Issuance of common stock in public offering, net of issuance costs (in shares) |  | 18,602,941 |  |  |
Issuance of common stock from exercise of stock options and units | 4,428 |  | 4,428 |  |
Issuance of common stock from exercise of stock options and units (in shares) |  | 599,529 |  |  |
Issuance of common stock pursuant to employee stock purchase plan | 3,682 |  | 3,682 |  |
Issuance of common stock pursuant to employee stock purchase plan (in shares) |  | 233,720 |  |  |
Issuance of common stock from exercise of warrants on a net issuance basis (in shares) |  | 6,855 |  |  |
Net loss | (245,192) |  |  | (245,192) |
Stock-based compensation | 82,312 |  | 82,312 |  |
Other comprehensive income (loss) | 27 |  |  |  | 27
Ending balance at Dec. 31, 2018 | 479,079 | $ 14 | 1,948,300 | (1,468,863) | (372)
Beginning balance (in shares) at Dec. 31, 2018 |  | 143,853,597 |  |  |
Issuance of common stock in public offering, net of issuance costs | $ 271,452 | $ 1 | 271,451 |  |
Issuance of common stock in public offering, net of issuance costs (in shares) |  | 7,187,500 |  |  |
Issuance of common stock from exercise of stock options (in shares) | 3,880,757 |  |  |  |
Issuance of common stock from exercise of stock options and units | $ 95,984 |  | 95,984 |  |
Issuance of common stock from exercise of stock options and units (in shares) |  | 3,965,166 |  |  |
Issuance of common stock pursuant to employee stock purchase plan | 4,411 |  | 4,411 |  |
Issuance of common stock pursuant to employee stock purchase plan (in shares) |  | 269,037 |  |  |
Net loss | (235,259) |  |  | (235,259) |
Stock-based compensation | 82,799 |  | 82,799 |  |
Other comprehensive income (loss) | 669 |  |  |  | 669
Ending balance at Dec. 31, 2019 | $ 699,135 | $ 15 | $ 2,402,945 | $ (1,704,122) | $ 297
Beginning balance (in shares) at Dec. 31, 2019 |  | 155,275,300 |  |  |

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