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Consolidated Statements of Stockholders' Equity - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Deficit
Accumulated Other Comprehensive Income (Loss)
Beginning balance at Dec. 31, 2016 $ 518,411 $ 12 $ 1,452,272 $ (933,979) $ 106
Beginning balance (in shares) at Dec. 31, 2016   121,367,169      
Issuance of common stock from exercise of stock options 26,665   26,665    
Issuance of common stock from exercise of stock options (in shares)   1,442,411      
Issuance of common stock pursuant to employee stock purchase plan 4,522   4,522    
Issuance of common stock pursuant to employee stock purchase plan (in shares)   192,402      
Issuance of common stock from exercise of warrants on a net issuance basis (in shares)   1,408,570      
Net loss (289,403)     (289,403)  
Cumulative effect adjustment from adoption of ASU 2016-09 (289)     (289)  
Stock-based compensation 75,884   75,884    
Other comprehensive income (loss) (505)       (505)
Ending balance at Dec. 31, 2017 335,285 $ 12 1,559,343 (1,223,671) (399)
Beginning balance (in shares) at Dec. 31, 2017   124,410,552      
Issuance of common stock in public offering, net of issuance costs 298,537 $ 2 298,535    
Issuance of common stock in public offering, net of issuance costs (in shares)   18,602,941      
Issuance of common stock from exercise of stock options and units 4,428   4,428    
Issuance of common stock from exercise of stock options and units (in shares)   599,529      
Issuance of common stock pursuant to employee stock purchase plan 3,682   3,682    
Issuance of common stock pursuant to employee stock purchase plan (in shares)   233,720      
Issuance of common stock from exercise of warrants on a net issuance basis (in shares)   6,855      
Net loss (245,192)     (245,192)  
Stock-based compensation 82,312   82,312    
Other comprehensive income (loss) 27       27
Ending balance at Dec. 31, 2018 479,079 $ 14 1,948,300 (1,468,863) (372)
Beginning balance (in shares) at Dec. 31, 2018   143,853,597      
Issuance of common stock in public offering, net of issuance costs $ 271,452 $ 1 271,451    
Issuance of common stock in public offering, net of issuance costs (in shares)   7,187,500      
Issuance of common stock from exercise of stock options (in shares) 3,880,757        
Issuance of common stock from exercise of stock options and units $ 95,984   95,984    
Issuance of common stock from exercise of stock options and units (in shares)   3,965,166      
Issuance of common stock pursuant to employee stock purchase plan 4,411   4,411    
Issuance of common stock pursuant to employee stock purchase plan (in shares)   269,037      
Net loss (235,259)     (235,259)  
Stock-based compensation 82,799   82,799    
Other comprehensive income (loss) 669       669
Ending balance at Dec. 31, 2019 $ 699,135 $ 15 $ 2,402,945 $ (1,704,122) $ 297
Beginning balance (in shares) at Dec. 31, 2019   155,275,300