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Business Combinations (Details) $ in Thousands |  | 12 Months Ended |
--- | --- | --- | ---
Jan. 08, 2016 USD ($) program platform | Dec. 31, 2016 USD ($) | Dec. 31, 2015 USD ($)
Business Acquisition [Line Items] |  |  |
Finite-lived intangible asset, useful life |  | 20 years |
Goodwill |  | $ 72,207 | $ 12,238
OMT, Inc. |  |  |
Business Acquisition [Line Items] |  |  |
Number of transgenic animal platforms | platform | 3 |  |
Number of partnered programs added | program | 16 |  |
Consideration transferred | $ 173,400 |  |
Goodwill | $ 59,969 |  |
Core Technology | OMT, Inc. |  |  |
Business Acquisition [Line Items] |  |  |
Projected cash flow discount rate (percent) | 15.50% |  |
Finite-lived intangible asset, useful life | 20 years |  |

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