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Acquisitions - Narrative (Details) |  |  |  |  | 3 Months Ended | 6 Months Ended
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Oct. 01, 2020 USD ($) | Sep. 09, 2020 USD ($) | Sep. 08, 2020 USD ($) | Apr. 01, 2020 USD ($) | Jun. 30, 2021 USD ($) | Dec. 31, 2020 USD ($) | Jun. 30, 2021 USD ($)
Pfenex |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Consideration transferred in acquisition | $ 465,100,000 |  |  |  |  |  |
Cash payments for acquisition | 429,600,000 |  |  |  |  |  |
Contingent earn-out payment | 77,800,000 |  |  |  |  |  |
Fair value of contingent earn-out payment | 37,000,000.0 |  |  |  |  |  |
Post-combination expense | 1,500,000 |  |  |  |  |  |
Gross payments to acquire business attributable to vested portion of stock options | $ 17,300,000 |  |  |  |  |  |
Cash to settle unvested stock options recognized as share-based compensation expense |  |  |  |  |  | $ 8,700,000 |
Weighted-average estimated useful life of finite-lived intangible assets acquired | 13 years |  |  |  |  |  |
Deferred tax liability acquired in connection with the acquired intangibles | $ (53,296,000) |  |  |  |  |  |
Deferred revenue acquired | (3,908,000) |  |  |  |  |  |
Pfenex | Contractual Relationships |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Finite-lived intangible assets acquired | $ 362,000,000 |  |  |  |  |  |
Weighted-average estimated useful life of finite-lived intangible assets acquired | 12 years 10 months 24 days |  |  |  |  |  |
Pfenex | Core Technology |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Finite-lived intangible assets acquired | $ 23,000,000 |  |  |  |  |  |
Pfenex | Core Technology | Minimum |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Discount rate used to value intangible assets acquired (as a percent) | 12.00% |  |  |  |  |  |
Pfenex | Core Technology | Maximum |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Discount rate used to value intangible assets acquired (as a percent) | 15.00% |  |  |  |  |  |
Pfenex | Discount Rate |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Contingent consideration liability, measurement input (as a percent) | 0.071 |  |  |  |  |  |
Taurus |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Consideration transferred in acquisition |  | $ 5,100,000 |  |  |  |  |
Cash payments for acquisition |  | 4,600,000 |  |  |  |  |
Contingent earn-out payment |  | 0 |  |  |  |  |
Holdback from acquisition |  | 500,000 |  |  |  |  |
Tangible assets acquired |  | 50,000.00 |  |  |  |  |
Taurus | Contingent Value Right for Internal Research and Development |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Contingent earn-out payment |  | 4,500,000 |  |  |  |  |
Taurus | Contingent Value Right on Product Revenues |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Contingent earn-out payment |  | $ 25,000,000.0 |  |  |  |  |
Proportion of milestone payments (as a percent) |  | 25.00% |  |  |  |  |
Taurus | Core Technology |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Finite-lived intangible assets acquired |  | $ 5,000,000.0 |  |  |  |  |
Weighted-average estimated useful life of finite-lived intangible assets acquired |  | 10 years |  |  |  |  |
xCella |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Consideration transferred in acquisition |  |  | $ 7,100,000 |  |  |  |
Contingent earn-out payment |  |  | 0 |  |  |  |
Holdback from acquisition |  |  | 500,000 |  |  |  |
Tangible assets acquired |  |  | 200,000 |  |  |  |
Liabilities assumed |  |  | (100,000) |  |  |  |
Deferred tax liability acquired in connection with the acquired intangibles |  |  | (800,000) |  |  |  |
xCella | Earnout Rights for Partner Research and Development |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Contingent earn-out payment |  |  | 5,000,000.0 |  |  |  |
Earnout rights recognized |  |  |  |  | $ 0 |  | $ 200,000
xCella | Milestone Payments |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Contingent earn-out payment |  |  | $ 25,750,000 |  |  |  |
Proportion of milestone payments (as a percent) |  |  | 25.00% |  |  |  |
xCella | Core Technology |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Finite-lived intangible assets acquired |  |  | $ 7,800,000 |  |  |  |
Weighted-average estimated useful life of finite-lived intangible assets acquired |  |  | 15 years |  |  |  |
Icagen |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Consideration transferred in acquisition |  |  |  | $ 19,900,000 |  |  |
Cash payments for acquisition |  |  |  | 15,100,000 |  |  |
Contingent earn-out payment |  |  |  | 25,000,000.0 |  |  |
Fair value of contingent earn-out payment |  |  |  | 4,800,000 |  |  |
Finite-lived intangible assets acquired |  |  |  | $ 12,800,000 |  |  |
Weighted-average estimated useful life of finite-lived intangible assets acquired |  |  |  | 9 years 8 months 12 days |  |  |
Tangible assets acquired |  |  |  | $ 1,800,000 |  |  |
Liabilities assumed |  |  |  | (800,000) |  |  |
Acquisition related costs |  |  |  | 0 |  |  |
Deferred revenue acquired |  |  |  | (3,700,000) |  |  |
Deferred tax assets acquired |  |  |  | 800,000 |  |  |
Goodwill acquired |  |  |  | 9,000,000.0 |  |  |
Icagen | Core Technology |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Finite-lived intangible assets acquired |  |  |  | $ 1,700,000 |  |  |
Weighted-average estimated useful life of finite-lived intangible assets acquired |  |  |  | 10 years |  |  |
Discount rate used to value intangible assets acquired (as a percent) |  |  |  | 17.00% |  |  |
Icagen | Customer Relationships |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |
Finite-lived intangible assets acquired |  |  |  | $ 11,100,000 |  |  |
Weighted-average estimated useful life of finite-lived intangible assets acquired |  |  |  | 9 years 7 months 6 days |  |  |
Discount rate used to value intangible assets acquired (as a percent) |  |  |  | 17.00% |  |  |

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