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Acquisitions - Narrative (Details) |  |  |  |  |  | 3 Months Ended | 12 Months Ended | 36 Months Ended
--- | --- | --- | --- | --- | --- | --- | --- | ---
Oct. 01, 2020 USD ($) | Sep. 09, 2020 USD ($) | Sep. 08, 2020 USD ($) | Apr. 01, 2020 USD ($) | Jul. 23, 2019 USD ($) | Dec. 31, 2020 USD ($) | Dec. 31, 2021 USD ($) | Dec. 31, 2020 USD ($) | Dec. 31, 2019 USD ($) | Dec. 31, 2021 acquisition
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Number of acquisitions | acquisition |  |  |  |  |  |  |  |  |  | 5
Number of business combinations | acquisition |  |  |  |  |  |  |  |  |  | 3
Number of asset acquisitions | acquisition |  |  |  |  |  |  |  |  |  | 2
Finite-lived intangible asset, useful life |  |  |  |  |  |  | 20 years |  |  |
Pfenex |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Consideration transferred | $ 465,100,000 |  |  |  |  |  |  |  |  |
Gross payments to acquire business | 429,600,000 |  |  |  |  |  |  |  |  |
Contingent consideration liability | 77,800,000 |  |  |  |  |  |  |  |  |
Fair value of contingent earn-out payment | 37,000,000 |  |  |  |  |  |  |  |  |
Post-combination expense | 1,500,000 |  |  |  |  |  |  |  |  |
Reduction in contingent liability |  |  |  |  |  |  | $ 37,600,000 |  |  |
Gross payments to acquire business attributable to vested portion of stock options | 17,300,000 |  |  |  |  |  |  |  |  |
Cash to settle unvested stock options recognized as share-based compensation expense |  |  |  |  |  | $ 8,700,000 |  |  |  |
Acquisition-related costs | $ 20,700,000 |  |  |  |  |  |  |  |  |
Revenue of acquiree since acquisition date included in comprehensive income |  |  |  |  |  |  |  | $ 2,000,000 |  |
Income (loss) before income taxes of acquiree since acquisition date included in comprehensive income |  |  |  |  |  |  |  | (19,300,000) |  |
Integration related costs |  |  |  |  |  |  |  | $ 7,100,000 | $ 4,800,000 |
Pfenex | Discount Rate |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Contingent consideration liability, measurement input (as a percent) | 0.071 |  |  |  |  |  |  |  |  |
Pfenex | Customer Relationships | Minimum |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Discount rate used to value intangible assets acquired (as a percent) |  |  |  | 12.00% |  |  |  |  |  |
Pfenex | Customer Relationships | Maximum |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Discount rate used to value intangible assets acquired (as a percent) |  |  |  | 15.00% |  |  |  |  |  |
Pfenex | Core Technology | Minimum |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Weighted-average useful lives of acquired finite-lived intangible assets | 10 years |  |  |  |  |  |  |  |  |
Pfenex | Core Technology | Maximum |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Weighted-average useful lives of acquired finite-lived intangible assets | 19 years |  |  |  |  |  |  |  |  |
Taurus |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Consideration transferred |  | $ 5,100,000 |  |  |  |  |  |  |  |
Gross payments to acquire business |  | 4,600,000 |  |  |  |  |  |  |  |
Contingent consideration liability |  | 0 |  |  |  |  |  |  |  |
Holdback from acquisition |  | 500,000 |  |  |  |  |  |  |  |
Acquired intangibles |  | 5,005,000 |  |  |  |  |  |  |  |
Taurus | Contingent Value Right for Internal Research and Development |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Contingent consideration liability |  | 4,500,000 |  |  |  |  |  |  |  |
Taurus | Contingent Value Right on Product Revenues |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Contingent consideration liability |  | $ 25,000,000 |  |  |  |  |  |  |  |
Proportion of milestone payments (as a percent) |  | 25.00% |  |  |  |  |  |  |  |
Taurus | Core Technology |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Weighted-average useful lives of acquired finite-lived intangible assets |  | 10 years |  |  |  |  |  |  |  |
xCella |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Consideration transferred |  |  | $ 7,100,000 |  |  |  |  |  |  |
Contingent consideration liability |  |  | 0 |  |  |  |  |  |  |
Holdback from acquisition |  |  | 500,000 |  |  |  |  |  |  |
Acquired intangibles |  |  | 7,582,000 |  |  |  |  |  |  |
xCella | Earnout Rights for Partner Research and Development |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Contingent consideration liability |  |  | 5,000,000 |  |  |  |  |  |  |
xCella | Milestone Payments |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Contingent consideration liability |  |  | $ 25,750,000 |  |  |  |  |  |  |
Proportion of milestone payments (as a percent) |  |  | 25.00% |  |  |  |  |  |  |
xCella | Core Technology |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Weighted-average useful lives of acquired finite-lived intangible assets |  |  | 15 years |  |  |  |  |  |  |
Icagen |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Consideration transferred |  |  |  | $ 19,900,000 |  |  |  |  |  |
Gross payments to acquire business |  |  |  | 15,100,000 |  |  |  |  |  |
Contingent consideration liability |  |  |  | 25,000,000 |  |  |  |  |  |
Fair value of contingent earn-out payment |  |  |  | $ 4,800,000 |  |  |  |  |  |
Weighted-average useful lives of acquired finite-lived intangible assets |  |  |  | 9 years 8 months 12 days |  |  |  |  |  |
Icagen | Discount Rate |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Contingent consideration liability, measurement input (as a percent) |  |  |  | 0.055 |  |  |  |  |  |
Icagen | Customer Relationships |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Discount rate used to value intangible assets acquired (as a percent) |  |  |  | 17.00% |  |  |  |  |  |
Weighted-average useful lives of acquired finite-lived intangible assets |  |  |  | 9 years 7 months 6 days |  |  |  |  |  |
Acquired intangibles |  |  |  | $ 11,100,000 |  |  |  |  |  |
Icagen | Core Technology |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Weighted-average useful lives of acquired finite-lived intangible assets |  |  |  | 10 years |  |  |  |  |  |
Acquired intangibles |  |  |  | $ 1,700,000 |  |  |  |  |  |
Ab Initio |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Consideration transferred |  |  |  |  | $ 12,000,000 |  |  |  |  |
Gross payments to acquire business |  |  |  |  | 11,900,000 |  |  |  |  |
Acquired intangibles |  |  |  |  | 7,400,000 |  |  |  |  |
Cash holdback for potential indemnification claims |  |  |  |  | 150,000 |  |  |  |  |
Goodwill acquired that is expected to be deductible for tax purposes |  |  |  |  | $ 0 |  |  |  |  |
Projected cash flow discount rate (as a percent) |  |  |  |  | 12.00% |  |  |  |  |
Ab Initio | Core Technology |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |
Finite-lived intangible asset, useful life |  |  |  |  | 20 years |  |  |  |  |

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