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Schedule III - Real Estate and Accumulated Depreciation (Details 2) - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2015 | Dec. 31, 2014 | Dec. 31, 2013
Changes in cost of properties |  |  |
Beginning balance | $ 4,014,336 | $ 3,811,950 | $ 3,859,960
Acquisitions of operating properties | 194,616 | 0 | 0
Building and land improvements | 273,761 | 254,868 | 249,639
Sales | (172,628) | (48,466) | (141,045)
Impairments | (29,548) | (3,042) | (45,931)
Other dispositions | (121,921) | (974) | (110,673)
Ending balance | 4,158,616 | 4,014,336 | 3,811,950
Changes in accumulated depreciation |  |  |
Beginning balance | 703,083 | 597,649 | 568,176
Depreciation expense | 112,695 | 111,326 | 92,677
Sales | (49,614) | (3,129) | (9,542)
Impairments | (6,092) | (1,671) | (14,863)
Other dispositions | (41,392) | (1,092) | (38,799)
Ending balance | $ 718,680 | $ 703,083 | $ 597,649

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- Definition Amount of accumulated depreciation pertaining to real estate investments for entities with a substantial portion of business acquiring and holding investment real estate. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 970 -SubTopic 360 -Section S99 -Paragraph 1 -Subparagraph (SX 210.12-28.FN4) -URI http://asc.fasb.org/extlink&oid=6590653&loc=d3e638233-123024 Reference 2: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 205 -SubTopic 10 -Section S99 -Paragraph 6 -Subparagraph (SX 210.5-04.(c) Schedule III) -URI http://asc.fasb.org/extlink&oid=27047687&loc=d3e5864-122674 + Details
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