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Income Taxes - Deferred Tax Assets (Liabilities) (Details) - USD ($) $ in Thousands | Dec. 31, 2021 | Dec. 31, 2020
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Deferred tax assets: |  |
Net operating loss carryforwards | $ 82,403 | $ 46,112
Tax credits | 25,800 | 17,423
Deferred revenue | 9,035 | 14,194
Operating lease liability | 7,365 | 9,196
Stock based compensation | 4,751 | 1,892
Reserve and accruals | 2,844 | 2,635
Total gross deferred tax assets | 132,198 | 91,452
Valuation allowance | (124,233) | (81,980)
Total deferred tax assets | 7,965 | 9,472
Operating lease right-of-use asset | (6,925) | (8,805)
Fixed and intangible assets | (1,040) | (667)
Total deferred tax liabilities | $ (7,965) | $ (9,472)

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