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Note 9 - Collaboration Agreements (Tables)
 9 Months Ended
Sep. 30, 2014
Organization, Consolidation and Presentation of Financial Statements [Abstract]  
Deferred Revenue, by Arrangement, Disclosure [Table Text Block]
    Three months ended
September 30,
 
 
Nine months ended
September 30,
 
    2013     2014     2013     2014  
                         
uniQure:
                       
Milestone payment
  $ 267     $ 14     $ 267     $ 14  
Teva:
                               
Recognition of upfront payment
    3,284       3,132       9,891       9,252  
Research funding
    169       84       463       251  
Genentech:
                               
Recognition of upfront payment
    824       981       2,483       2,736  
Research funding
    1,128       994       3,385       3,249  
       Milestone payment
    5,093       7,987       5,093       7,987  
Genome BC:
                               
Research funding
    21       -       189       -  
Total collaboration revenue
  $ 10,786     $ 13,192     $ 21,771     $ 23,489