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Cash, Cash Equivalents, Marketable Securities, and Restricted Cash - Narrative (Details) - USD ($)
12 Months Ended
Dec. 31, 2021
Dec. 31, 2020
Dec. 31, 2019
Cash, Cash Equivalents, and Short-term Investments [Abstract]      
Cash and cash equivalents $ 245,197,000 $ 163,240,000 $ 142,837,000
Debt securities, available-for-sale 237,299,000 320,029,000  
Debt securities, realized gain (loss) 0 0  
Fair value of available-for-sale debt securities in unrealized loss positions $ 173,400,000 $ 124,900,000