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Leases
 12 Months Ended
Dec. 31, 2024
Leases [Abstract]  
Leases Leases
The Company currently has operating leases for office and research laboratory space, equipment, and vehicles under agreements expiring at various dates through 2034, which include renewal options on leases which the Company is reasonably certain to exercise.
For the years ended December 31, 2024 and 2023, operating lease expense was $6.7 million and $9.5 million and variable lease expense was $2.2 million and $2.1 million, respectively. For the years ended December 31, 2024 and 2023, the Company paid $9.6 million and $9.5 million, respectively, for amounts included in the measurement of operating lease liabilities and recorded right-of-use assets of $0.7 million and $0.6 million, respectively.
Commitments under finance leases are not significant for the year ended December 31, 2024.
Supplemental balance sheet information related to operating leases were as follows:
December 31,
(in thousands, except year and discount rate amounts)20242023
Operating lease ROU assets, net$22,278 $26,312 
Current portion of the operating lease liabilities$8,455 $8,324 
Non-current portion of the operating lease liabilities45,078 48,877 
Total operating lease liability$53,533 $57,201 
Weighted-average remaining lease terms (years)8.59.4
Weighted-average discount rate9.7 %9.7 %
At December 31, 2024, the future minimum operating lease payments were as follows:
(in thousands)Operating Lease
2025$9,466 
20269,356 
20279,426 
20289,139 
20298,172 
Thereafter35,402 
Total lease payments80,961 
Less: lease incentives(22,299)
Less: imputed interest(5,129)
Total operating lease liability$53,533