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Summary of Significant Accounting Policies - Additional Information (Detail) - USD ($)
$ in Thousands
1 Months Ended 3 Months Ended
Feb. 28, 2010
Mar. 31, 2018
Accounting Policies [Line Items]    
Variable consideration related to milestone payment   $ 0
Revenue recognized for milestone payment   $ 0
Intrexon Corporation    
Accounting Policies [Line Items]    
Nonrefundable upfront payment $ 400