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Condensed Consolidated Statements of Cash Flows (unaudited) - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2018
Mar. 31, 2017
Operating activities:    
Net loss $ (16,749) $ (13,398)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 139 198
Stock-based compensation 1,461 1,051
Amortization of debt issuance costs and debt discount 20 19
Amortization of discount or premium on short-term investments 36 (39)
Deferred rent (10) (5)
Changes in assets and liabilities:    
Prepaid expenses and other current assets 482 (416)
Accounts payable and accrued liabilities 549 (799)
Accrued compensation (650) (1,129)
Net cash used in operating activities (14,722) (14,518)
Investing activities:    
Purchases of short-term investments (5,803) 0
Maturities of short-term investments 6,285 11,400
Purchases of property and equipment (31) (116)
Net cash provided by investing activities 451 11,284
Financing activities:    
Proceeds from issuance of common stock, net of issuance costs 6,440 0
Proceeds from exercise of stock options 28 144
Repurchase of unvested restricted stock 0 (79)
Net cash provided by financing activities 6,468 65
Net decrease in cash, cash equivalents, and restricted cash (7,803) (3,169)
Cash, cash equivalents, and restricted cash at beginning of period 60,813 85,367
Cash, cash equivalents, and restricted cash at end of period 53,010 82,198
Supplemental disclosure of cash flows:    
Interest paid 137 118
Non-cash financing activities:    
Vesting of early exercised stock options 9 16
Deferred financing costs incurred but not yet paid $ 133 $ 0