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Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-in Capital | Accumulated Deficit
--- | --- | --- | --- | ---
Balance at Dec. 31, 2013 | $ 56,443 | $ 21 | $ 142,071 | $ (85,649)
Balance (in shares) at Dec. 31, 2013 |  | 21,143,313 |  |
Increase (Decrease) in Convertible Preferred Stock and Stockholders' Equity (Deficit) |  |  |  |
Issuance of common stock on exercise of stock options | 675 | $ 1 | 674 |
Issuance of common stock on exercise of stock options (in shares) |  | 402,100 |  |
Issuance of common stock on cashless exercise of warrant (in shares) |  | 13,739 |  |
Common stock subject to repurchase | 3 |  | 3 |
Issuance of common stock for acquisition | 10,078 | $ 1 | 10,077 |
Issuance of common stock for acquisition (in shares) |  | 964,377 |  |
Stock-based compensation expense (employee) | 3,388 |  | 3,388 |
Stock-based compensation expense (non-employee) | 160 |  | 160 |
Net loss | (29,373) |  |  | (29,373)
Comprehensive loss | (29,373) |  |  | (29,373)
Balance at Dec. 31, 2014 | 41,374 | $ 23 | 156,373 | (115,022)
Balance (in shares) at Dec. 31, 2014 |  | 22,523,529 |  |
Increase (Decrease) in Convertible Preferred Stock and Stockholders' Equity (Deficit) |  |  |  |
Issuance of common stock on exercise of stock options | 722 |  | 722 |
Issuance of common stock on exercise of stock options (in shares) |  | 253,787 |  |
Issuance of common stock for acquisition | 0 |  |  |
Sale of common stock in a private placement, net of issuance costs | 37,258 | $ 5 | 37,253 |
Sale of common stock in a private placement, net of issuance costs (in shares) |  | 4,907,975 |  |
Issuance of common stock in a private placement, issuance cost | 2,742 |  |  |
Stock-based compensation expense (employee) | 5,302 |  | 5,302 |
Stock-based compensation expense (non-employee) | 110 |  | 110 |
Stock-based compensation expense (ESPP) | 190 |  | 190 |
Net loss | (33,704) |  |  | (33,704)
Comprehensive loss | (33,704) |  |  | (33,704)
Balance at Dec. 31, 2015 | 51,252 | $ 28 | 199,950 | (148,726)
Balance (in shares) at Dec. 31, 2015 |  | 27,685,291 |  |
Increase (Decrease) in Convertible Preferred Stock and Stockholders' Equity (Deficit) |  |  |  |
Issuance of common stock on exercise of stock options | $ 538 |  | 538 |
Issuance of common stock on exercise of stock options (in shares) | 212,740 | 212,740 |  |
Issuance of common stock under employee stock purchase plan (ESPP) | $ 678 |  | 678 |
Issuance of common stock on employee stock purchase plan (ESPP) (in shares) |  | 140,947 |  |
Issuance of common stock for acquisition | 0 |  |  |
Sale of common stock in a private placement, net of issuance costs | 32,093 | $ 6 | 32,087 |
Sale of common stock in a private placement, net of issuance costs (in shares) |  | 5,723,300 |  |
Issuance of common stock in a private placement, issuance cost | 2,247 |  |  |
Stock-based compensation expense (employee) | 6,046 |  | 6,046 |
Stock-based compensation expense (non-employee) | 15 |  | 15 |
Stock-based compensation expense (ESPP) | 317 |  | 317 |
Net loss | (31,358) |  |  | (31,358)
Comprehensive loss | (31,358) |  |  | (31,358)
Balance at Dec. 31, 2016 | $ 59,581 | $ 34 | $ 239,631 | $ (180,084)
Balance (in shares) at Dec. 31, 2016 |  | 33,762,278.000 |  |

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