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Balance Sheet Components (Tables)
 12 Months Ended
Dec. 31, 2016
Balance Sheet Related Disclosures [Abstract]  
Schedule of property and equipment
Property and equipment consisted of the following (in thousands of dollars):

 
Year Ended December 31,
 
2016
 
2015
Leasehold improvements
$
5,861

 
$
789

Laboratory equipment
6,441

 
5,501

Computer equipment
1,177

 
1,046

Software, including software developed for internal use
1,937

 
1,353

Furniture and fixtures
1,131

 
242

Construction-in-process
1,769

 
6,823

Total property and equipment, at cost
18,316

 
15,754

Accumulated depreciation and amortization
(6,836
)
 
(5,440
)
Total property and equipment, net
$
11,480

 
$
10,314
Schedule of accrued liabilities
Accrued liabilities consisted of the following (in thousands of dollars):

 
Year Ended December 31,
 
2016
 
2015
Accrued compensation expense
$
6,120

 
$
4,212

Accrued Genzyme co-promotion fees
—

 
2,089

Accrued other
2,990

 
2,388

Total accrued liabilities
$
9,110

 
$
8,689