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Balance Sheet Components - Property and Equipment, Net (Details) - USD ($)
$ in Thousands
 12 Months Ended
Dec. 31, 2016
Dec. 31, 2015
Dec. 31, 2014
Property and Equipment, Net      
Total property and equipment, at cost $ 18,316 $ 15,754  
Accumulated depreciation and amortization (6,836) (5,440)  
Total property and equipment, net 11,480 10,314  
Depreciation and amortization expense 3,511 2,254 $ 1,175
Depreciation and amortization expense 2,400 1,500  
Leasehold improvements      
Property and Equipment, Net      
Total property and equipment, at cost 5,861 789  
Laboratory equipment      
Property and Equipment, Net      
Total property and equipment, at cost 6,441 5,501  
Computer equipment      
Property and Equipment, Net      
Total property and equipment, at cost 1,177 1,046  
Software, including software developed for internal use      
Property and Equipment, Net      
Total property and equipment, at cost 1,937 1,353  
Furniture and fixtures      
Property and Equipment, Net      
Total property and equipment, at cost 1,131 242  
Construction-in-process      
Property and Equipment, Net      
Total property and equipment, at cost $ 1,769 $ 6,823