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Collaboration Agreements - Biogen (Details) | 3 Months Ended | 9 Months Ended |
--- | --- | --- | ---
Sep. 30, 2021 USD ($) milestone | Sep. 30, 2020 USD ($) | Sep. 30, 2021 USD ($) milestone | Sep. 30, 2020 USD ($) | Dec. 31, 2020 USD ($)
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Offset to research and development expense, related party cost reimbursement | $ 1,200,000 |  |  | $ 5,300,000 |  |  |
Cost sharing reimbursements due from related party | 1,194,000 |  |  | 1,194,000 |  |  | $ 5,674,000
Collaboration revenue from customers | 5,285,000 | [1] | $ 9,388,000 | 36,143,000 | [1] | $ 18,751,000 |
Biogen Collaboration Agreement | Related Party |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Contract liability | 294,900,000 |  |  | 294,900,000 |  |  |
Biogen Collaborative Arrangement | Biogen Collaboration Agreement |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Transaction price, change | 0 |  |  | 0 |  |  |
Offset to research and development expense, related party cost reimbursement | 1,200,000 |  |  | 5,300,000 |  |  |
Cost sharing reimbursements due from related party | $ 1,200,000 |  |  | $ 1,200,000 |  |  | $ 5,700,000
Number of milestones achieved | milestone | 0 |  |  | 0 |  |  |
Biogen Collaborative Arrangement | Biogen Collaboration Agreement | Option Research Services | Related Party |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Contract liability | $ 6,000,000.0 |  |  | $ 6,000,000.0 |  |  |
Biogen Collaborative Arrangement | Biogen Collaboration Agreement | Product |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Collaboration revenue from customers |  |  |  | 0 |  |  |
ROFN and Option Agreement | Biogen Collaboration Agreement | Related Party |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Contract liability | $ 288,900,000 |  |  | $ 288,900,000 |  |  |
[1] | Includes related party collaboration revenue from a customer of $0.9 million and $2.5 million for the three and nine months ended September 30, 2021, respectively.

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