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Collaboration Agreements (Tables) | 9 Months Ended
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Sep. 30, 2021
Organization, Consolidation and Presentation of Financial Statements [Abstract] |
Summary of Collaboration Revenue | Revenue disaggregated by collaboration agreement and performance obligation is as follows (in thousands):
| Three Months Ended September 30, |  | Nine Months Ended September 30,
| 2021 |  | 2020 |  | 2021 |  | 2020
Takeda Collaboration Agreement(1) | $ | 4,425 |  |  | $ | 9,271 |  |  | $ | 18,597 |  |  | $ | 17,826 |
Sanofi Collaboration Agreement |  |  |  |  |  |  |
Peripheral Program License | — |  |  | — |  |  | 15,000 |  |  | — |
Retained Activities | — |  |  | 117 |  |  | — |  |  | 925 |
Alzheimer's Disease Services(2) | — |  |  | 5 |  |  | 4 |  |  | 93 |
Total Sanofi Collaboration Revenue | — |  |  | 122 |  |  | 15,004 |  |  | 1,018 |
Biogen Collaboration Agreement |  |  |  |  |  |  |
Option Research Services(2) | 860 |  |  | — |  |  | 2,546 |  |  | — |
Total Biogen Collaboration Revenue | $ | 860 |  |  | $ | — |  |  | $ | 2,546 |  |  | $ | — |
Total Collaboration Revenue | $ | 5,285 |  |  | $ | 9,393 |  |  | $ | 36,147 |  |  | $ | 18,844 |
_________________________________________________ (1)$15.6 million of revenue for the nine months ended September 30, 2021 was included in the contract liability balance at the beginning of the period. All revenue for the three months ended September 30, 2021 and the three and nine months ended September 30, 2020 was included in the contract liability balance at the beginning of the period. (2)Revenue for the three and nine months ended September 30, 2021 and September 30, 2020, where applicable, represent amounts that were included in the contract liability balance at the beginning of the respective period.

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- Definition Tabular disclosure of disaggregation of revenue into categories depicting how nature, amount, timing, and uncertainty of revenue and cash flows are affected by economic factor. + References Reference 1: http://www.xbrl.org/2009/role/commonPracticeRef -Publisher FASB -Name Accounting Standards Codification -Topic 606 -SubTopic 10 -Section 50 -Paragraph 5 -URI http://asc.fasb.org/extlink&oid=123351226&loc=SL49130545-203045 + Details
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