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Collaboration Agreements - Sanofi (Details) - USD ($)	1 Months Ended	3 Months Ended			9 Months Ended			36 Months Ended
	Jun. 30, 2021	Sep. 30, 2021		Sep. 30, 2020	Sep. 30, 2021		Sep. 30, 2020	Sep. 30, 2021	Dec. 31, 2020
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Collaboration revenue from customers		$ 5,285,000	[1]	$ 9,388,000	$ 36,143,000	[1]	$ 18,751,000
Sanofi | Collaborative Arrangement
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Receivable		0			0			$ 0	$ 44,303
Sanofi | Collaborative Arrangement | Milestone Triggered
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Transaction price increase		0			15,000,000
Collaboration revenue from customers	$ 15,000,000							25,000,000
Sanofi | Collaborative Arrangement | Milestone Triggered and Retained Activities
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Transaction price increase				$ 100,000			$ 900,000
Sanofi | Collaborative Arrangement | Alzheimer's Disease Services
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Contract liability		$ 3,400,000			$ 3,400,000			3,400,000	$ 3,400,000
Sanofi | Collaborative Arrangement | Product
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Collaboration revenue from customers								$ 0
[1] Includes related party collaboration revenue from a customer of $0.9 million and $2.5 million for the three and nine months ended September 30, 2021, respectively.

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