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Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($)
$ in Thousands
 3 Months Ended 9 Months Ended
Sep. 30, 2021
Sep. 30, 2020
Sep. 30, 2021
Sep. 30, 2020
Collaboration revenue:        
Collaboration revenue from customers $ 5,285 [1] $ 9,388 $ 36,143 [1] $ 18,751
Other collaboration revenue 0 5 4 93
Total collaboration revenue 5,285 9,393 36,147 18,844
Operating expenses:        
Research and development 71,559 [2] 53,704 197,477 [2] 157,872
General and administrative 19,319 15,805 57,300 42,332
Total operating expenses 90,878 69,509 254,777 200,204
Loss from operations (85,593) (60,116) (218,630) (181,360)
Interest and other income, net 1,005 1,944 3,310 7,611
Loss before income taxes (84,588) (58,172) (215,320) (173,749)
Income tax expense 0 (56) 0 0
Net loss (84,588) (58,228) (215,320) (173,749)
Other comprehensive loss:        
Net unrealized loss on marketable securities, net of tax (153) (540) (276) (339)
Comprehensive loss $ (84,741) $ (58,768) $ (215,596) $ (174,088)
Net loss per share, basic (usd per share) $ (0.69) $ (0.54) $ (1.77) $ (1.65)
Net loss per share, diluted (usd per share) $ (0.69) $ (0.54) $ (1.77) $ (1.65)
Weighted average number of shares outstanding, basic (in shares) 121,742,067 107,490,702 121,309,197 105,217,770
Weighted average number of shares outstanding, diluted (in shares) 121,742,067 107,490,702 121,309,197 105,217,770
[1] Includes related party collaboration revenue from a customer of $0.9 million and $2.5 million for the three and nine months ended September 30, 2021, respectively.
[2] Includes an offset to expense from related party cost reimbursement of $1.2 million and $5.3 million for the three and nine months ended September 30, 2021, respectively.