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Consolidated Statements of Shareholders' Equity - USD ($) $ in Thousands | Total | Revision of Prior Period, Accounting Standards Update, Adjustment | Common Stock | Additional Paid In Capital | Accumulated Other Comprehensive Gain (Loss) | Accumulated Deficit | Accumulated Deficit Revision of Prior Period, Accounting Standards Update, Adjustment | Noncontrolling Interest
--- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning Balance at Dec. 31, 2019 | $ 571,039 | $ 93 | $ 12 | $ 935,834 | $ 671 | $ (365,478) | $ 93 |
Beginning Balance, Shares at Dec. 31, 2019 |  |  | 125,238,142 |  |  |  |  |
Issuance of common stock upon public offering, after deducting underwriters' discounts and net offering costs paid by us | $ 271,839 |  | $ 1 | 271,838 |  |  |  |
Issuance of common stock upon public offering, after deducting underwriters' discounts and net offering costs paid by us, Shares |  |  | 7,200,000 |  |  |  |  |
Accounting Standards Update Extensible List | ASC 842 |  |  |  |  |  |  |
Issuance of common stock for cash upon exercise of stock options | $ 21,539 |  | $ 1 | 21,538 |  |  |  |
Issuance of common stock for cash upon exercise of stock options, Shares |  |  | 5,204,254 |  |  |  |  |
Vesting of restricted stock units |  |  | 4,500 |  |  |  |  |
Share-based compensation | 24,761 |  |  | 24,761 |  |  |  |
Other comprehensive income (loss) | 222 |  |  |  | 222 |  |  |
Net loss | (146,227) |  |  |  |  | (146,227) |  |
Ending Balance at Dec. 31, 2020 | 743,266 |  | $ 14 | 1,253,971 | 893 | (511,612) |  |
Ending Balance, Shares at Dec. 31, 2020 |  |  | 137,646,896 |  |  |  |  |
Issuance of common stock upon exercise of common stock warrant, Shares |  |  | 54,162 |  |  |  |  |
Issuance of common stock for cash upon exercise of stock options | 26,484 |  |  | 26,484 |  |  |  |
Issuance of common stock for cash upon exercise of stock options, Shares |  |  | 3,674,057 |  |  |  |  |
Vesting of restricted stock units |  |  | 18,750 |  |  |  |  |
Share-based compensation | 43,251 |  |  | 43,251 |  |  |  |
Capital contributions for Digital Biotechnologies, Inc. | 429 |  |  | 300 |  |  |  | $ 129
Other comprehensive income (loss) | (2,030) |  |  |  | (2,030) |  |  |
Net loss | (207,298) |  |  |  |  | (207,279) |  | (19)
Ending Balance at Dec. 31, 2021 | 604,102 |  | $ 14 | 1,324,006 | (1,137) | (718,891) |  | 110
Ending Balance, Shares at Dec. 31, 2021 |  |  | 141,393,865 |  |  |  |  |
Issuance of common stock for cash upon exercise of stock options | $ 7,866 |  |  | 7,866 |  |  |  |
Issuance of common stock for cash upon exercise of stock options, Shares | 1,406,500 |  | 1,406,500 |  |  |  |  |
Vesting of restricted stock units |  |  | 304,637 |  |  |  |  |
Share-based compensation | $ 55,477 |  |  | 55,477 |  |  |  |
Other comprehensive income (loss) | (2,979) |  |  |  | (2,979) |  |  |
Net loss | (200,368) |  |  |  |  | (200,191) |  | (177)
Ending Balance at Dec. 31, 2022 | $ 464,098 |  | $ 14 | $ 1,387,349 | $ (4,116) | $ (919,082) |  | $ (67)
Ending Balance, Shares at Dec. 31, 2022 |  |  | 143,105,002 |  |  |  |  |

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