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Income Taxes - Summary of Deferred tax assets and liabilities (Details) - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2019 | Dec. 31, 2018 | Dec. 31, 2017
Gross Deferred Tax Assets |  |  |
Net Operating Loss Carryforwards | $ 172,745 | $ 168,239 |
Foreign Net Operating Loss Carryforwards |  | 4,485 |
Tax Credit Carryforwards | 42,642 | 39,761 |
Deferred Research and Development Expenses | 70,042 | 76,555 |
Stock-based Compensation | 12,651 | 11,977 |
Fixed Assets | 1,759 | 1,761 |
Accrued Expenses and Other | 328 | 316 |
Deferred Tax Assets, Gross | 300,167 | 303,094 |
Gross Deferred Tax Liabilities |  |  |
IPR&D Intangibles | (12,748) | (12,748) |
Total Deferred Tax Assets and Liabilities | 287,419 | 290,346 |
Valuation Allowance | (290,426) | (293,353) |
Net Deferred Tax Liability | $ 3,007 | $ 3,007 |
Principal components of the deferred tax assets and liabilities |  |  |
Increase in deferred tax and valuation allowance |  |  | $ 17,700
U.S. federal corporate income tax rate (as a percent) | 21.00% | 21.00% | 35.00%
U.S. federal corporate income tax net operating losses (as a percent) |  |  | 80.00%
Change in tax rate, deferred tax liability benefit |  | $ 6,900 |
Additional income tax benefit leading to tax liabilities |  | 12,200 |
Revaluation of deferred tax asset |  | 99,500 |
Income tax expense (benefit) |  | $ (765) | $ (24,282)

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