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Income Taxes - Summary of Deferred tax assets and liabilities (Details) - USD ($) $ in Thousands | 12 Months Ended |
--- | --- | ---
Dec. 31, 2020 | Dec. 31, 2018 | Dec. 31, 2019
Gross deferred tax assets |  |  |
Net operating loss carryforwards | $ 174,369 |  | $ 172,745
Tax credit carryforwards | 44,936 |  | 42,642
Deferred research and development expenses | 63,215 |  | 70,042
Stock-based compensation | 13,196 |  | 12,651
Fixed assets | 1,303 |  | 1,759
Accrued expenses and other | 325 |  | 328
Gross deferred tax assets | 297,344 |  | 300,167
Gross deferred tax liabilities |  |  |
IPR&D intangibles | 7,802 |  | 12,748
Total deferred tax assets and liabilities | 289,542 |  | 287,419
Valuation Allowance | (291,382) |  | (290,426)
Net deferred tax liability | 1,840 |  | $ 3,007
Principal components of the deferred tax assets and liabilities |  |  |
Income Tax Benefit | $ (1,167) | $ (765) |

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