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CONDENSED CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($) shares in Thousands, $ in Thousands | 3 Months Ended | 9 Months Ended
--- | --- | ---
Sep. 30, 2021 | Sep. 30, 2020 | Sep. 30, 2021 | Sep. 30, 2020
Revenues: |  |  |  |
Total revenues | $ 153 | $ 668 | $ 4,317 | $ 3,633
Operating expenses: |  |  |  |
Research and development | 13,557 | 10,708 | 38,633 | 32,109
General and administrative | 5,821 | 3,640 | 14,247 | 10,833
Intangible asset impairment | 3,500 |  | 3,500 | 3,500
(Gain) loss on fair value remeasurement of contingent consideration | (1,901) | 662 | (1,160) | (4,236)
Total operating expenses | 20,977 | 15,010 | 55,220 | 42,206
Operating loss | (20,824) | (14,342) | (50,903) | (38,573)
Investment and other income, net | 145 | 118 | 313 | 465
Net loss before income tax benefit | (20,679) | (14,224) | (50,590) | (38,108)
Income tax benefit | 227 |  | 227 | 228
Net loss | $ (20,452) | $ (14,224) | $ (50,363) | $ (37,880)
Basic and diluted net loss per common share | $ (0.45) | $ (0.36) | $ (1.21) | $ (1.44)
Shares used in calculating basic and diluted net loss per share | 45,453 | 39,278 | 41,582 | 26,303
Comprehensive loss: |  |  |  |
Net loss | $ (20,452) | $ (14,224) | $ (50,363) | $ (37,880)
Other comprehensive income (loss): |  |  |  |
Unrealized (loss) gain on marketable securities | (44) | 14 | (41) | (11)
Comprehensive loss | (20,496) | (14,210) | (50,404) | (37,891)
Product development and licensing agreements |  |  |  |
Revenues: |  |  |  |
Total revenues |  | 12 | 29 | 2,297
Contracts and grants |  |  |  |
Revenues: |  |  |  |
Total revenues | $ 153 | $ 656 | $ 4,288 | $ 1,336

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