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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2021 | Dec. 31, 2020 | Dec. 31, 2019
Revenues: |  |  |
Total revenues | $ 4,651 | $ 7,418 | $ 3,573
Operating expenses: |  |  |
Research and development | 53,311 | 42,534 | 42,672
General and administrative | 20,488 | 14,456 | 15,426
Intangible asset impairment | 3,500 | 18,000 |
Other asset impairment |  |  | 1,800
Gain on fair value remeasurement of contingent consideration | (1,405) | (4,218) | (1,294)
Total operating expenses | 75,894 | 70,772 | 58,604
Operating loss | (71,243) | (63,354) | (55,031)
Investment and other income, net | 505 | 2,407 | 4,153
Net loss before income tax benefit | (70,738) | (60,947) | (50,878)
Income tax benefit | 227 | 1,167 |
Net loss | $ (70,511) | $ (59,780) | $ (50,878)
Basic net loss per common share | $ (1.64) | $ (2.02) | $ (3.51)
Diluted net loss per common share | $ (1.64) | $ (2.02) | $ (3.51)
Shares used in calculating basic net loss per share | 42,870 | 29,640 | 14,507
Shares used in calculating diluted net loss per share | 42,870 | 29,640 | 14,507
Comprehensive loss: |  |  |
Net loss | $ (70,511) | $ (59,780) | $ (50,878)
Other comprehensive income (loss): |  |  |
Unrealized (loss) gain on marketable securities | (695) | (30) | 36
Comprehensive loss | (71,206) | (59,810) | (50,842)
Product development and licensing agreements |  |  |
Revenues: |  |  |
Total revenues | 31 | 2,301 | 473
Contracts and grants |  |  |
Revenues: |  |  |
Total revenues | $ 4,620 | $ 5,117 | $ 3,100

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