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Income Taxes - Summary of Deferred tax assets and liabilities (Details) - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2021 | Dec. 31, 2020
Gross deferred tax assets |  |
Net operating loss carryforwards | $ 183,711 | $ 174,369
Tax credit carryforwards | 47,927 | 44,936
Deferred research and development expenses | 59,753 | 63,215
Stock-based compensation | 14,719 | 13,196
Fixed assets | 1,287 | 1,303
Accrued expenses and other | 370 | 325
Gross deferred tax assets | 307,767 | 297,344
Gross deferred tax liabilities |  |
IPR&D intangibles | (6,840) | (7,802)
Total deferred tax assets and liabilities | 300,927 | 289,542
Valuation Allowance | (302,540) | (291,382)
Net deferred tax liability | 1,613 | 1,840
Principal components of the deferred tax assets and liabilities |  |
Income Tax Benefit | (227) | (1,167)
IPR&D |  |
Principal components of the deferred tax assets and liabilities |  |
Income Tax Benefit | $ 200 | $ 1,200

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