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Stockholders' Equity (Details) - USD ($) $ in Thousands | 3 Months Ended | 9 Months Ended
--- | --- | ---
Sep. 30, 2023 | Jun. 30, 2023 | Mar. 31, 2023 | Sep. 30, 2022 | Jun. 30, 2022 | Mar. 31, 2022 | Sep. 30, 2023 | Sep. 30, 2022
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Balance, beginning | $ 277,537 | $ 302,740 | $ 326,204 | $ 365,093 | $ 398,101 | $ 419,476 | $ 326,204 | $ 419,476
Balance, beginning (in shares) |  |  | 47,200,695 |  |  |  | 47,200,695 |
Shares issued under stock option and employee stock purchase plans | 257 | 133 | $ 694 | 2,306 | 71 | 304 |  |
Stock-based compensation | 7,121 | 5,217 | 4,340 | 4,496 | 3,454 | 3,153 |  |
Unrealized gain (loss) on marketable securities | 63 | (51) | 863 | 305 | (529) | (1,782) | $ 875 | (2,006)
Net loss | (38,260) | (30,502) | (29,361) | (26,784) | (36,004) | (23,050) | (98,123) | (85,838)
Balance, ending | $ 246,718 | 277,537 | 302,740 | 345,416 | 365,093 | 398,101 | $ 246,718 | 345,416
Balance, ending (in shares) | 47,264,197 |  |  |  |  |  | 47,264,197 |
Aggregate gross offering price available | $ 50,000 |  |  |  |  |  | $ 50,000 |
Common Stock |  |  |  |  |  |  |  |
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Balance, beginning | $ 47 | $ 47 | $ 47 | $ 47 | $ 47 | $ 47 | $ 47 | $ 47
Balance, beginning (in shares) | 47,253,813 | 47,244,681 | 47,200,695 | 46,764,703 | 46,754,348 | 46,730,198 | 47,200,695 | 46,730,198
Shares issued under stock option and employee stock purchase plans (in shares) | 10,384 | 9,132 | 43,986 | 331,360 | 10,355 | 24,150 |  |
Balance, ending | $ 47 | $ 47 | $ 47 | $ 47 | $ 47 | $ 47 | $ 47 | $ 47
Balance, ending (in shares) | 47,264,197 | 47,253,813 | 47,244,681 | 47,096,063 | 46,764,703 | 46,754,348 | 47,264,197 | 47,096,063
Additional Paid-In Capital |  |  |  |  |  |  |  |
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Balance, beginning | $ 1,591,213 | $ 1,585,863 | $ 1,580,829 | $ 1,568,124 | $ 1,564,599 | $ 1,561,142 | $ 1,580,829 | $ 1,561,142
Shares issued under stock option and employee stock purchase plans | 257 | 133 | 694 | 2,306 | 71 | 304 |  |
Stock-based compensation | 7,121 | 5,217 | 4,340 | 4,496 | 3,454 | 3,153 |  |
Balance, ending | 1,598,591 | 1,591,213 | 1,585,863 | 1,574,926 | 1,568,124 | 1,564,599 | 1,598,591 | 1,574,926
Accumulated Other Comprehensive Income |  |  |  |  |  |  |  |
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Balance, beginning | 2,072 | 2,123 | 1,260 | (417) | 112 | 1,894 | 1,260 | 1,894
Unrealized gain (loss) on marketable securities | 63 | (51) | 863 | 305 | (529) | (1,782) |  |
Balance, ending | 2,135 | 2,072 | 2,123 | (112) | (417) | 112 | 2,135 | (112)
Accumulated Deficit |  |  |  |  |  |  |  |
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Balance, beginning | (1,315,795) | (1,285,293) | (1,255,932) | (1,202,661) | (1,166,657) | (1,143,607) | (1,255,932) | (1,143,607)
Net loss | (38,260) | (30,502) | (29,361) | (26,784) | (36,004) | (23,050) |  |
Balance, ending | $ (1,354,055) | $ (1,315,795) | $ (1,285,293) | $ (1,229,445) | $ (1,202,661) | $ (1,166,657) | $ (1,354,055) | $ (1,229,445)

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