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Consolidated Statements of Stockholders' Equity - USD ($) shares in Thousands, $ in Thousands | Common Stock [Member] | Additional Paid-in Capital [Member] | Accumulated Deficit [Member] | Accumulated Other Comprehensive Income (Loss) [Member] | Total
--- | --- | --- | --- | --- | ---
Beginning Balance at Dec. 31, 2014 | $ 28 | $ 271,282 | $ (138,424) | $ (348) | $ 132,538
Beginning Balance, Shares at Dec. 31, 2014 | 27,580 |  |  |  |
Issuance of common stock through public offering | $ 3 | 68,985 |  |  | 68,988
Issuance of common stock through public offering, Shares | 3,757 |  |  |  |
Issuance of common stock under equity incentive plans | $ 1 | 1,920 |  |  | 1,921
Issuance of common stock under equity incentive plans, Shares | 850 |  |  |  |
Issuance of common stock under employee stock purchase plan |  | 1,539 |  |  | 1,539
Issuance of common stock under employee stock purchase plan, Shares | 87 |  |  |  |
Reclassification of non-employee option liability |  | 177 |  |  | 177
Share-based employee compensation expense |  | 8,997 |  |  | 8,997
Other comprehensive loss |  |  |  | (263) | (263)
Net loss |  |  | (27,212) |  | (27,212)
Ending Balance at Dec. 31, 2015 | $ 32 | 352,900 | (165,636) | (611) | 186,685
Ending Balance, Shares at Dec. 31, 2015 | 32,274 |  |  |  |
Issuance of common stock under equity incentive plans | $ 1 | 1,636 |  |  | 1,637
Issuance of common stock under equity incentive plans, Shares | 934 |  |  |  |
Issuance of common stock under employee stock purchase plan |  | 1,618 |  |  | 1,618
Issuance of common stock under employee stock purchase plan, Shares | 134 |  |  |  |
Share-based employee compensation expense |  | 11,697 |  |  | 11,697
Other comprehensive loss |  |  |  | 143 | 143
Net loss |  |  | (33,338) |  | (33,338)
Ending Balance at Dec. 31, 2016 | $ 33 | 367,851 | (198,974) | (468) | 168,442
Ending Balance, Shares at Dec. 31, 2016 | 33,342 |  |  |  |
Issuance of common stock under equity incentive plans | $ 2 | 2,387 |  |  | 2,389
Issuance of common stock under equity incentive plans, Shares | 1,112 |  |  |  |
Issuance of common stock under employee stock purchase plan |  | 2,110 |  |  | 2,110
Issuance of common stock under employee stock purchase plan, Shares | 132 |  |  |  |
Share-based employee compensation expense |  | 14,615 |  |  | 14,615
Other comprehensive loss |  |  |  | 502 | 502
Net loss |  |  | (26,892) |  | (26,892)
Ending Balance at Dec. 31, 2017 | $ 35 | $ 386,963 | $ (225,866) | $ 34 | $ 161,166
Ending Balance, Shares at Dec. 31, 2017 | 34,586 |  |  |  |

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