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Consolidated Statements of Stockholders' Equity - USD ($)
shares in Thousands, $ in Thousands
Common Stock [Member]
Additional Paid-in Capital [Member]
Accumulated Deficit [Member]
Accumulated Other Comprehensive Income (Loss) [Member]
Total
Beginning Balance at Dec. 31, 2014 $ 28 $ 271,282 $ (138,424) $ (348) $ 132,538
Beginning Balance, Shares at Dec. 31, 2014 27,580        
Issuance of common stock through public offering $ 3 68,985 68,988
Issuance of common stock through public offering, Shares 3,757        
Issuance of common stock under equity incentive plans $ 1 1,920 1,921
Issuance of common stock under equity incentive plans, Shares 850        
Issuance of common stock under employee stock purchase plan 1,539 1,539
Issuance of common stock under employee stock purchase plan, Shares 87        
Reclassification of non-employee option liability 177 177
Share-based employee compensation expense 8,997 8,997
Other comprehensive loss (263) (263)
Net loss (27,212) (27,212)
Ending Balance at Dec. 31, 2015 $ 32 352,900 (165,636) (611) 186,685
Ending Balance, Shares at Dec. 31, 2015 32,274        
Issuance of common stock under equity incentive plans $ 1 1,636 1,637
Issuance of common stock under equity incentive plans, Shares 934        
Issuance of common stock under employee stock purchase plan 1,618 1,618
Issuance of common stock under employee stock purchase plan, Shares 134        
Share-based employee compensation expense 11,697 11,697
Other comprehensive loss 143 143
Net loss (33,338) (33,338)
Ending Balance at Dec. 31, 2016 $ 33 367,851 (198,974) (468) 168,442
Ending Balance, Shares at Dec. 31, 2016 33,342        
Issuance of common stock under equity incentive plans $ 2 2,387 2,389
Issuance of common stock under equity incentive plans, Shares 1,112        
Issuance of common stock under employee stock purchase plan 2,110 2,110
Issuance of common stock under employee stock purchase plan, Shares 132        
Share-based employee compensation expense 14,615 14,615
Other comprehensive loss 502 502
Net loss (26,892) (26,892)
Ending Balance at Dec. 31, 2017 $ 35 $ 386,963 $ (225,866) $ 34 $ 161,166
Ending Balance, Shares at Dec. 31, 2017 34,586