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Condensed Consolidated Statements of Stockholders' Deficit - USD ($) shares in Thousands, $ in Thousands | Total | Senior Convertible Notes | Facility Financing Obligation | Related Party Notes | At-the-market Issuance | Common Stock | Common Stock Senior Convertible Notes | Common Stock Facility Financing Obligation | Common Stock Related Party Notes | Common Stock At-the-market Issuance | Additional Paid-in Capital | Additional Paid-in Capital Senior Convertible Notes | Additional Paid-in Capital Facility Financing Obligation | Additional Paid-in Capital Related Party Notes | Additional Paid-in Capital At-the-market Issuance | Accumulated Other Comprehensive Income (Loss) | Accumulated Deficit
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning Balance at Dec. 31, 2017 | $ (214,733) |  |  |  |  | $ 1,191 |  |  |  |  | $ 2,638,992 |  |  |  |  | $ (18) | $ (2,854,898)
Beginning Balance (in shares) at Dec. 31, 2017 |  |  |  |  |  | 119,053 |  |  |  |  |  |  |  |  |  |  |
Adjustment to adopt ASU 2016-09 | Topic 718 | 1,873 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 1,873
Issuance of common shares from the release of restricted stock units | (81) |  |  |  |  | $ 1 |  |  |  |  | (82) |  |  |  |  |  |
Issuance of common shares from the release of restricted stock units (in shares) |  |  |  |  |  | 60 |  |  |  |  |  |  |  |  |  |  |
Issuance of common shares under Employee Stock Purchase Plan | 1 |  |  |  |  | $ 1 |  |  |  |  |  |  |  |  |  |  |
Issuance of common shares under Employee Stock Purchase Plan (in shares) |  |  |  |  |  | 137 |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense | 1,943 |  |  |  |  |  |  |  |  |  | 1,943 |  |  |  |  |  |
Issuance of shares pursuant to conversion notes |  |  | $ 9,407 | $ 8,160 |  |  |  | $ 35 | $ 30 |  |  |  | $ 9,372 | $ 8,130 |  |  |
Issuance of shares pursuant to conversion notes (in shares) |  |  |  |  |  |  |  | 3,549 | 3,000 |  |  |  |  |  |  |  |
Cumulative translation gain | 3 |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 3 |
Amortization of shelf fees | (5) |  |  |  |  |  |  |  |  |  | (5) |  |  |  |  |  |
Issuance of at-the-market placement |  |  |  |  | $ 634 |  |  |  |  | $ 2 |  |  |  |  | $ 632 |  |
Issuance of at-the-market placement (in shares) |  |  |  |  |  |  |  |  |  | 214 |  |  |  |  |  |  |
Issuance costs associated with at-the- market placement | (25) |  |  |  |  |  |  |  |  |  | (25) |  |  |  |  |  |
Net loss | (30,385) |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (30,385)
Ending Balance at Mar. 31, 2018 | (223,208) |  |  |  |  | $ 1,260 |  |  |  |  | 2,658,957 |  |  |  |  | (15) | (2,883,410)
Ending Balance (in shares) at Mar. 31, 2018 |  |  |  |  |  | 126,013 |  |  |  |  |  |  |  |  |  |  |
Beginning Balance at Dec. 31, 2018 | (175,082) |  |  |  |  | $ 1,870 |  |  |  |  | 2,763,067 |  |  |  |  | (19) | (2,940,000)
Beginning Balance (in shares) at Dec. 31, 2018 |  |  |  |  |  | 187,030 |  |  |  |  |  |  |  |  |  |  |
Exercise of stock options | 3 |  |  |  |  |  |  |  |  |  | 3 |  |  |  |  |  |
Exercise of stock options (in shares) |  |  |  |  |  | 3 |  |  |  |  |  |  |  |  |  |  |
Issuance of common shares from the release of restricted stock units | (1) |  |  |  |  | $ 1 |  |  |  |  | (2) |  |  |  |  |  |
Issuance of common shares from the release of restricted stock units (in shares) |  |  |  |  |  | 63 |  |  |  |  |  |  |  |  |  |  |
Issuance of common shares under Employee Stock Purchase Plan | 317 |  |  |  |  | $ 3 |  |  |  |  | 314 |  |  |  |  |  |
Issuance of common shares under Employee Stock Purchase Plan (in shares) |  |  |  |  |  | 296 |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense | 999 |  |  |  |  |  |  |  |  |  | 999 |  |  |  |  |  |
Restricted stock unit award | 105 |  |  |  |  |  |  |  |  |  | 105 |  |  |  |  |  |
Issuance of shares pursuant to conversion notes |  | $ 538 |  |  |  |  | $ 4 |  |  |  |  | $ 534 |  |  |  |  |
Issuance of shares pursuant to conversion notes (in shares) |  |  |  |  |  |  | 386 |  |  |  |  |  |  |  |  |  |
Net loss | (14,883) |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (14,883)
Ending Balance at Mar. 31, 2019 | $ (188,004) |  |  |  |  | $ 1,878 |  |  |  |  | $ 2,765,020 |  |  |  |  | $ (19) | $ (2,954,883)
Ending Balance (in shares) at Mar. 31, 2019 |  |  |  |  |  | 187,778 |  |  |  |  |  |  |  |  |  |  |

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